Amino acid dipepetide frequency for Anabas testudineus (Climbing perch) (Anthias testudineus)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	5.638AlaAla: 5.638 ± 0.021	1.258AlaCys: 1.258 ± 0.008	3.09AlaAsp: 3.09 ± 0.01	4.415AlaGlu: 4.415 ± 0.018	2.413AlaPhe: 2.413 ± 0.01	4.024AlaGly: 4.024 ± 0.013	1.426AlaHis: 1.426 ± 0.007	2.897AlaIle: 2.897 ± 0.009	3.379AlaLys: 3.379 ± 0.012	6.447AlaLeu: 6.447 ± 0.018	1.6AlaMet: 1.6 ± 0.007	2.239AlaAsn: 2.239 ± 0.008	3.075AlaPro: 3.075 ± 0.013	2.812AlaGln: 2.812 ± 0.011	2.999AlaArg: 2.999 ± 0.01	4.994AlaSer: 4.994 ± 0.013	3.392AlaThr: 3.392 ± 0.01	4.837AlaVal: 4.837 ± 0.011	0.658AlaTrp: 0.658 ± 0.004	1.603AlaTyr: 1.603 ± 0.006	0.0AlaXaa: 0.0 ± 0.0
Cys	1.182CysAla: 1.182 ± 0.006	0.654CysCys: 0.654 ± 0.005	1.181CysAsp: 1.181 ± 0.01	1.289CysGlu: 1.289 ± 0.01	0.998CysPhe: 0.998 ± 0.005	1.514CysGly: 1.514 ± 0.009	0.679CysHis: 0.679 ± 0.005	1.106CysIle: 1.106 ± 0.007	1.185CysLys: 1.185 ± 0.008	2.313CysLeu: 2.313 ± 0.012	0.49CysMet: 0.49 ± 0.004	0.883CysAsn: 0.883 ± 0.006	1.275CysPro: 1.275 ± 0.009	1.074CysGln: 1.074 ± 0.009	1.3CysArg: 1.3 ± 0.008	2.123CysSer: 2.123 ± 0.01	1.236CysThr: 1.236 ± 0.006	1.698CysVal: 1.698 ± 0.009	0.31CysTrp: 0.31 ± 0.003	0.668CysTyr: 0.668 ± 0.004	0.0CysXaa: 0.0 ± 0.0
Asp	2.885AspAla: 2.885 ± 0.01	1.199AspCys: 1.199 ± 0.007	3.068AspAsp: 3.068 ± 0.014	3.714AspGlu: 3.714 ± 0.013	2.24AspPhe: 2.24 ± 0.008	3.586AspGly: 3.586 ± 0.013	1.218AspHis: 1.218 ± 0.005	2.91AspIle: 2.91 ± 0.011	2.852AspLys: 2.852 ± 0.01	5.089AspLeu: 5.089 ± 0.013	1.332AspMet: 1.332 ± 0.005	2.102AspAsn: 2.102 ± 0.009	2.849AspPro: 2.849 ± 0.011	1.98AspGln: 1.98 ± 0.008	2.699AspArg: 2.699 ± 0.011	4.29AspSer: 4.29 ± 0.013	2.709AspThr: 2.709 ± 0.009	3.354AspVal: 3.354 ± 0.011	0.704AspTrp: 0.704 ± 0.005	1.687AspTyr: 1.687 ± 0.008	0.0AspXaa: 0.0 ± 0.0
Glu	4.546GluAla: 4.546 ± 0.018	1.298GluCys: 1.298 ± 0.01	4.265GluAsp: 4.265 ± 0.014	7.176GluGlu: 7.176 ± 0.028	2.059GluPhe: 2.059 ± 0.007	4.041GluGly: 4.041 ± 0.012	1.487GluHis: 1.487 ± 0.007	3.025GluIle: 3.025 ± 0.009	4.642GluLys: 4.642 ± 0.016	6.315GluLeu: 6.315 ± 0.021	1.759GluMet: 1.759 ± 0.006	2.81GluAsn: 2.81 ± 0.009	2.608GluPro: 2.608 ± 0.01	3.09GluGln: 3.09 ± 0.013	4.108GluArg: 4.108 ± 0.016	4.137GluSer: 4.137 ± 0.011	3.375GluThr: 3.375 ± 0.011	4.355GluVal: 4.355 ± 0.012	0.723GluTrp: 0.723 ± 0.005	1.724GluTyr: 1.724 ± 0.007	0.0GluXaa: 0.0 ± 0.0
Phe	2.014PheAla: 2.014 ± 0.008	1.017PheCys: 1.017 ± 0.005	1.899PheAsp: 1.899 ± 0.008	1.982PheGlu: 1.982 ± 0.008	1.782PhePhe: 1.782 ± 0.009	2.311PheGly: 2.311 ± 0.01	1.068PheHis: 1.068 ± 0.005	2.171PheIle: 2.171 ± 0.01	1.919PheLys: 1.919 ± 0.008	4.12PheLeu: 4.12 ± 0.013	0.874PheMet: 0.874 ± 0.005	1.619PheAsn: 1.619 ± 0.006	1.836PhePro: 1.836 ± 0.008	1.68PheGln: 1.68 ± 0.007	1.903PheArg: 1.903 ± 0.008	3.45PheSer: 3.45 ± 0.011	2.467PheThr: 2.467 ± 0.009	2.419PheVal: 2.419 ± 0.01	0.5PheTrp: 0.5 ± 0.004	1.341PheTyr: 1.341 ± 0.007	0.0PheXaa: 0.0 ± 0.0
Gly	3.809GlyAla: 3.809 ± 0.015	1.225GlyCys: 1.225 ± 0.007	3.198GlyAsp: 3.198 ± 0.01	3.981GlyGlu: 3.981 ± 0.013	2.464GlyPhe: 2.464 ± 0.01	4.931GlyGly: 4.931 ± 0.022	1.666GlyHis: 1.666 ± 0.014	2.777GlyIle: 2.777 ± 0.01	3.624GlyLys: 3.624 ± 0.011	5.417GlyLeu: 5.417 ± 0.015	1.46GlyMet: 1.46 ± 0.007	2.511GlyAsn: 2.511 ± 0.009	3.027GlyPro: 3.027 ± 0.022	2.714GlyGln: 2.714 ± 0.01	3.455GlyArg: 3.455 ± 0.013	5.364GlySer: 5.364 ± 0.018	3.479GlyThr: 3.479 ± 0.018	4.001GlyVal: 4.001 ± 0.012	0.785GlyTrp: 0.785 ± 0.005	1.935GlyTyr: 1.935 ± 0.01	0.0GlyXaa: 0.0 ± 0.0
His	1.29HisAla: 1.29 ± 0.006	0.81HisCys: 0.81 ± 0.006	1.02HisAsp: 1.02 ± 0.005	1.267HisGlu: 1.267 ± 0.007	1.118HisPhe: 1.118 ± 0.005	1.542HisGly: 1.542 ± 0.007	0.996HisHis: 0.996 ± 0.007	1.443HisIle: 1.443 ± 0.007	1.327HisLys: 1.327 ± 0.006	2.718HisLeu: 2.718 ± 0.01	0.657HisMet: 0.657 ± 0.004	1.087HisAsn: 1.087 ± 0.006	1.547HisPro: 1.547 ± 0.013	1.225HisGln: 1.225 ± 0.007	1.591HisArg: 1.591 ± 0.006	2.378HisSer: 2.378 ± 0.009	1.64HisThr: 1.64 ± 0.007	1.493HisVal: 1.493 ± 0.007	0.348HisTrp: 0.348 ± 0.003	0.912HisTyr: 0.912 ± 0.005	0.0HisXaa: 0.0 ± 0.0
Ile	2.709IleAla: 2.709 ± 0.01	1.166IleCys: 1.166 ± 0.007	2.321IleAsp: 2.321 ± 0.008	2.569IleGlu: 2.569 ± 0.01	2.026IlePhe: 2.026 ± 0.01	2.445IleGly: 2.445 ± 0.009	1.367IleHis: 1.367 ± 0.005	2.725IleIle: 2.725 ± 0.012	2.777IleLys: 2.777 ± 0.008	4.677IleLeu: 4.677 ± 0.015	1.176IleMet: 1.176 ± 0.005	2.209IleAsn: 2.209 ± 0.008	2.527IlePro: 2.527 ± 0.008	2.332IleGln: 2.332 ± 0.009	2.598IleArg: 2.598 ± 0.009	3.924IleSer: 3.924 ± 0.011	2.999IleThr: 2.999 ± 0.009	2.812IleVal: 2.812 ± 0.011	0.514IleTrp: 0.514 ± 0.004	1.588IleTyr: 1.588 ± 0.008	0.0IleXaa: 0.0 ± 0.0
Lys	3.703LysAla: 3.703 ± 0.013	1.068LysCys: 1.068 ± 0.006	3.336LysAsp: 3.336 ± 0.012	4.719LysGlu: 4.719 ± 0.016	1.741LysPhe: 1.741 ± 0.008	3.205LysGly: 3.205 ± 0.012	1.475LysHis: 1.475 ± 0.007	2.692LysIle: 2.692 ± 0.01	4.404LysLys: 4.404 ± 0.018	5.279LysLeu: 5.279 ± 0.014	1.476LysMet: 1.476 ± 0.007	2.427LysAsn: 2.427 ± 0.009	2.74LysPro: 2.74 ± 0.01	2.603LysGln: 2.603 ± 0.01	3.257LysArg: 3.257 ± 0.011	3.724LysSer: 3.724 ± 0.012	3.266LysThr: 3.266 ± 0.01	3.68LysVal: 3.68 ± 0.012	0.603LysTrp: 0.603 ± 0.003	1.621LysTyr: 1.621 ± 0.008	0.0LysXaa: 0.0 ± 0.0
Leu	6.052LeuAla: 6.052 ± 0.017	2.351LeuCys: 2.351 ± 0.009	5.05LeuAsp: 5.05 ± 0.013	6.511LeuGlu: 6.511 ± 0.023	3.743LeuPhe: 3.743 ± 0.014	5.138LeuGly: 5.138 ± 0.013	2.783LeuHis: 2.783 ± 0.009	4.166LeuIle: 4.166 ± 0.012	5.798LeuLys: 5.798 ± 0.016	10.28LeuLeu: 10.28 ± 0.027	2.219LeuMet: 2.219 ± 0.008	3.86LeuAsn: 3.86 ± 0.01	5.066LeuPro: 5.066 ± 0.014	5.495LeuGln: 5.495 ± 0.019	5.585LeuArg: 5.585 ± 0.015	8.274LeuSer: 8.274 ± 0.022	5.455LeuThr: 5.455 ± 0.013	5.68LeuVal: 5.68 ± 0.014	1.094LeuTrp: 1.094 ± 0.006	2.832LeuTyr: 2.832 ± 0.01	0.0LeuXaa: 0.0 ± 0.0
Met	1.849MetAla: 1.849 ± 0.007	0.523MetCys: 0.523 ± 0.004	1.449MetAsp: 1.449 ± 0.006	1.981MetGlu: 1.981 ± 0.007	0.935MetPhe: 0.935 ± 0.005	1.414MetGly: 1.414 ± 0.006	0.507MetHis: 0.507 ± 0.004	0.967MetIle: 0.967 ± 0.005	1.515MetLys: 1.515 ± 0.006	2.2MetLeu: 2.2 ± 0.008	0.702MetMet: 0.702 ± 0.005	0.958MetAsn: 0.958 ± 0.005	1.049MetPro: 1.049 ± 0.006	1.016MetGln: 1.016 ± 0.005	1.181MetArg: 1.181 ± 0.006	1.85MetSer: 1.85 ± 0.007	1.304MetThr: 1.304 ± 0.006	1.631MetVal: 1.631 ± 0.007	0.283MetTrp: 0.283 ± 0.003	0.695MetTyr: 0.695 ± 0.004	0.0MetXaa: 0.0 ± 0.0
Asn	2.176AsnAla: 2.176 ± 0.008	0.954AsnCys: 0.954 ± 0.007	1.813AsnAsp: 1.813 ± 0.008	2.176AsnGlu: 2.176 ± 0.008	1.533AsnPhe: 1.533 ± 0.006	2.786AsnGly: 2.786 ± 0.013	1.041AsnHis: 1.041 ± 0.006	2.401AsnIle: 2.401 ± 0.009	2.365AsnLys: 2.365 ± 0.007	3.868AsnLeu: 3.868 ± 0.013	1.101AsnMet: 1.101 ± 0.006	1.953AsnAsn: 1.953 ± 0.008	2.291AsnPro: 2.291 ± 0.009	1.784AsnGln: 1.784 ± 0.008	2.035AsnArg: 2.035 ± 0.007	3.282AsnSer: 3.282 ± 0.012	2.418AsnThr: 2.418 ± 0.009	2.439AsnVal: 2.439 ± 0.009	0.478AsnTrp: 0.478 ± 0.004	1.261AsnTyr: 1.261 ± 0.007	0.0AsnXaa: 0.0 ± 0.0
Pro	3.639ProAla: 3.639 ± 0.012	1.049ProCys: 1.049 ± 0.007	2.832ProAsp: 2.832 ± 0.011	3.584ProGlu: 3.584 ± 0.011	1.763ProPhe: 1.763 ± 0.007	3.859ProGly: 3.859 ± 0.036	1.449ProHis: 1.449 ± 0.007	1.948ProIle: 1.948 ± 0.007	2.405ProLys: 2.405 ± 0.013	4.662ProLeu: 4.662 ± 0.013	0.995ProMet: 0.995 ± 0.006	1.938ProAsn: 1.938 ± 0.007	4.725ProPro: 4.725 ± 0.027	2.466ProGln: 2.466 ± 0.012	2.476ProArg: 2.476 ± 0.009	4.999ProSer: 4.999 ± 0.02	2.933ProThr: 2.933 ± 0.014	3.655ProVal: 3.655 ± 0.011	0.544ProTrp: 0.544 ± 0.004	1.437ProTyr: 1.437 ± 0.007	0.0ProXaa: 0.0 ± 0.0
Gln	3.104GlnAla: 3.104 ± 0.013	1.063GlnCys: 1.063 ± 0.009	2.323GlnAsp: 2.323 ± 0.008	3.403GlnGlu: 3.403 ± 0.013	1.488GlnPhe: 1.488 ± 0.006	2.698GlnGly: 2.698 ± 0.011	1.352GlnHis: 1.352 ± 0.007	2.088GlnIle: 2.088 ± 0.009	2.597GlnLys: 2.597 ± 0.01	4.59GlnLeu: 4.59 ± 0.014	1.145GlnMet: 1.145 ± 0.006	1.777GlnAsn: 1.777 ± 0.007	2.37GlnPro: 2.37 ± 0.012	3.153GlnGln: 3.153 ± 0.022	2.952GlnArg: 2.952 ± 0.011	3.327GlnSer: 3.327 ± 0.013	2.538GlnThr: 2.538 ± 0.01	2.9GlnVal: 2.9 ± 0.009	0.575GlnTrp: 0.575 ± 0.004	1.351GlnTyr: 1.351 ± 0.006	0.0GlnXaa: 0.0 ± 0.0
Arg	3.239ArgAla: 3.239 ± 0.01	1.262ArgCys: 1.262 ± 0.008	2.901ArgAsp: 2.901 ± 0.011	3.767ArgGlu: 3.767 ± 0.012	1.999ArgPhe: 1.999 ± 0.008	3.272ArgGly: 3.272 ± 0.013	1.5ArgHis: 1.5 ± 0.006	2.505ArgIle: 2.505 ± 0.008	3.437ArgLys: 3.437 ± 0.01	5.303ArgLeu: 5.303 ± 0.014	1.299ArgMet: 1.299 ± 0.006	2.191ArgAsn: 2.191 ± 0.007	2.683ArgPro: 2.683 ± 0.011	2.653ArgGln: 2.653 ± 0.01	3.933ArgArg: 3.933 ± 0.014	4.126ArgSer: 4.126 ± 0.013	2.86ArgThr: 2.86 ± 0.008	3.327ArgVal: 3.327 ± 0.009	0.679ArgTrp: 0.679 ± 0.005	1.614ArgTyr: 1.614 ± 0.007	0.0ArgXaa: 0.0 ± 0.0
Ser	5.163SerAla: 5.163 ± 0.015	1.977SerCys: 1.977 ± 0.01	4.223SerAsp: 4.223 ± 0.013	4.643SerGlu: 4.643 ± 0.012	3.153SerPhe: 3.153 ± 0.009	5.355SerGly: 5.355 ± 0.018	2.173SerHis: 2.173 ± 0.009	3.501SerIle: 3.501 ± 0.01	3.916SerLys: 3.916 ± 0.013	8.15SerLeu: 8.15 ± 0.022	1.795SerMet: 1.795 ± 0.006	3.036SerAsn: 3.036 ± 0.01	5.291SerPro: 5.291 ± 0.022	3.708SerGln: 3.708 ± 0.012	4.196SerArg: 4.196 ± 0.011	9.261SerSer: 9.261 ± 0.035	4.755SerThr: 4.755 ± 0.015	5.416SerVal: 5.416 ± 0.013	0.972SerTrp: 0.972 ± 0.006	2.225SerTyr: 2.225 ± 0.008	0.0SerXaa: 0.0 ± 0.0
Thr	3.876ThrAla: 3.876 ± 0.011	1.432ThrCys: 1.432 ± 0.011	2.99ThrAsp: 2.99 ± 0.01	3.756ThrGlu: 3.756 ± 0.013	2.258ThrPhe: 2.258 ± 0.01	3.762ThrGly: 3.762 ± 0.013	1.469ThrHis: 1.469 ± 0.007	2.622ThrIle: 2.622 ± 0.01	2.759ThrLys: 2.759 ± 0.01	5.565ThrLeu: 5.565 ± 0.013	1.257ThrMet: 1.257 ± 0.006	2.043ThrAsn: 2.043 ± 0.008	3.451ThrPro: 3.451 ± 0.017	2.37ThrGln: 2.37 ± 0.009	2.458ThrArg: 2.458 ± 0.008	4.702ThrSer: 4.702 ± 0.013	3.443ThrThr: 3.443 ± 0.032	4.377ThrVal: 4.377 ± 0.013	0.694ThrTrp: 0.694 ± 0.005	1.571ThrTyr: 1.571 ± 0.007	0.0ThrXaa: 0.0 ± 0.0
Val	4.156ValAla: 4.156 ± 0.011	1.812ValCys: 1.812 ± 0.011	3.326ValAsp: 3.326 ± 0.01	4.117ValGlu: 4.117 ± 0.011	2.841ValPhe: 2.841 ± 0.01	3.568ValGly: 3.568 ± 0.012	1.636ValHis: 1.636 ± 0.006	3.247ValIle: 3.247 ± 0.01	3.733ValLys: 3.733 ± 0.011	6.374ValLeu: 6.374 ± 0.015	1.592ValMet: 1.592 ± 0.007	2.64ValAsn: 2.64 ± 0.009	3.195ValPro: 3.195 ± 0.011	2.81ValGln: 2.81 ± 0.009	3.319ValArg: 3.319 ± 0.01	5.332ValSer: 5.332 ± 0.013	4.062ValThr: 4.062 ± 0.011	4.557ValVal: 4.557 ± 0.014	0.813ValTrp: 0.813 ± 0.005	1.948ValTyr: 1.948 ± 0.008	0.0ValXaa: 0.0 ± 0.0
Trp	0.675TrpAla: 0.675 ± 0.004	0.269TrpCys: 0.269 ± 0.002	0.672TrpAsp: 0.672 ± 0.005	0.774TrpGlu: 0.774 ± 0.004	0.479TrpPhe: 0.479 ± 0.004	0.626TrpGly: 0.626 ± 0.005	0.284TrpHis: 0.284 ± 0.003	0.61TrpIle: 0.61 ± 0.005	0.764TrpLys: 0.764 ± 0.005	1.184TrpLeu: 1.184 ± 0.006	0.357TrpMet: 0.357 ± 0.003	0.548TrpAsn: 0.548 ± 0.004	0.43TrpPro: 0.43 ± 0.003	0.514TrpGln: 0.514 ± 0.004	0.733TrpArg: 0.733 ± 0.004	0.967TrpSer: 0.967 ± 0.006	0.731TrpThr: 0.731 ± 0.005	0.718TrpVal: 0.718 ± 0.005	0.212TrpTrp: 0.212 ± 0.002	0.365TrpTyr: 0.365 ± 0.003	0.0TrpXaa: 0.0 ± 0.0
Tyr	1.494TyrAla: 1.494 ± 0.006	0.773TyrCys: 0.773 ± 0.005	1.512TyrAsp: 1.512 ± 0.007	1.716TyrGlu: 1.716 ± 0.007	1.338TyrPhe: 1.338 ± 0.007	1.746TyrGly: 1.746 ± 0.008	0.832TyrHis: 0.832 ± 0.005	1.661TyrIle: 1.661 ± 0.008	1.623TyrLys: 1.623 ± 0.008	2.881TyrLeu: 2.881 ± 0.012	0.724TyrMet: 0.724 ± 0.004	1.324TyrAsn: 1.324 ± 0.005	1.319TyrPro: 1.319 ± 0.008	1.315TyrGln: 1.315 ± 0.007	1.775TyrArg: 1.775 ± 0.008	2.402TyrSer: 2.402 ± 0.009	1.792TyrThr: 1.792 ± 0.007	1.726TyrVal: 1.726 ± 0.007	0.426TyrTrp: 0.426 ± 0.005	1.102TyrTyr: 1.102 ± 0.007	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.0XaaAla: 0.0 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.0XaaAsp: 0.0 ± 0.0	0.0XaaGlu: 0.0 ± 0.0	0.0XaaPhe: 0.0 ± 0.0	0.0XaaGly: 0.0 ± 0.0	0.0XaaHis: 0.0 ± 0.0	0.0XaaIle: 0.0 ± 0.0	0.0XaaLys: 0.0 ± 0.0	0.0XaaLeu: 0.0 ± 0.0	0.0XaaMet: 0.0 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.0XaaPro: 0.0 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.0XaaArg: 0.0 ± 0.0	0.0XaaSer: 0.0 ± 0.0	0.0XaaThr: 0.0 ± 0.0	0.0XaaVal: 0.0 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.002XaaXaa: 0.002 ± 0.001

Statistics based on 61942 proteins (42338991 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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