Amino acid dipepetide frequency for Tortoise microvirus 85

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
3.199AlaAla: 3.199 ± 2.523
0.0AlaCys: 0.0 ± 0.0
1.919AlaAsp: 1.919 ± 0.862
8.317AlaGlu: 8.317 ± 3.023
2.559AlaPhe: 2.559 ± 1.381
8.957AlaGly: 8.957 ± 3.516
0.0AlaHis: 0.0 ± 0.0
7.038AlaIle: 7.038 ± 1.861
3.839AlaLys: 3.839 ± 1.138
2.559AlaLeu: 2.559 ± 1.262
2.559AlaMet: 2.559 ± 1.801
4.479AlaAsn: 4.479 ± 1.632
3.199AlaPro: 3.199 ± 1.614
3.839AlaGln: 3.839 ± 1.245
3.199AlaArg: 3.199 ± 1.325
1.28AlaSer: 1.28 ± 1.097
3.199AlaThr: 3.199 ± 1.267
1.919AlaVal: 1.919 ± 1.275
0.64AlaTrp: 0.64 ± 0.416
2.559AlaTyr: 2.559 ± 1.117
0.0AlaXaa: 0.0 ± 0.0
Cys
0.64CysAla: 0.64 ± 0.627
0.0CysCys: 0.0 ± 0.0
0.0CysAsp: 0.0 ± 0.0
0.0CysGlu: 0.0 ± 0.0
0.64CysPhe: 0.64 ± 0.627
0.0CysGly: 0.0 ± 0.0
0.0CysHis: 0.0 ± 0.0
0.64CysIle: 0.64 ± 0.416
0.0CysLys: 0.0 ± 0.0
0.0CysLeu: 0.0 ± 0.0
0.0CysMet: 0.0 ± 0.0
0.0CysAsn: 0.0 ± 0.0
0.0CysPro: 0.0 ± 0.0
0.0CysGln: 0.0 ± 0.0
0.64CysArg: 0.64 ± 0.627
0.0CysSer: 0.0 ± 0.0
0.0CysThr: 0.0 ± 0.0
0.64CysVal: 0.64 ± 0.416
0.0CysTrp: 0.0 ± 0.0
0.0CysTyr: 0.0 ± 0.0
0.0CysXaa: 0.0 ± 0.0
Asp
3.839AspAla: 3.839 ± 2.351
0.0AspCys: 0.0 ± 0.0
3.839AspAsp: 3.839 ± 1.999
3.839AspGlu: 3.839 ± 2.522
0.64AspPhe: 0.64 ± 0.627
3.199AspGly: 3.199 ± 1.99
2.559AspHis: 2.559 ± 2.194
4.479AspIle: 4.479 ± 1.672
4.479AspLys: 4.479 ± 1.31
5.758AspLeu: 5.758 ± 2.405
2.559AspMet: 2.559 ± 1.004
2.559AspAsn: 2.559 ± 0.861
1.28AspPro: 1.28 ± 0.832
3.199AspGln: 3.199 ± 0.939
2.559AspArg: 2.559 ± 1.198
1.28AspSer: 1.28 ± 0.559
3.199AspThr: 3.199 ± 1.466
3.839AspVal: 3.839 ± 1.172
3.199AspTrp: 3.199 ± 0.966
1.919AspTyr: 1.919 ± 1.267
0.0AspXaa: 0.0 ± 0.0
Glu
6.398GluAla: 6.398 ± 2.865
0.64GluCys: 0.64 ± 0.627
3.199GluAsp: 3.199 ± 1.837
6.398GluGlu: 6.398 ± 5.063
0.0GluPhe: 0.0 ± 0.0
4.479GluGly: 4.479 ± 1.55
1.919GluHis: 1.919 ± 1.502
7.038GluIle: 7.038 ± 1.818
9.597GluLys: 9.597 ± 2.984
5.118GluLeu: 5.118 ± 3.12
0.64GluMet: 0.64 ± 0.416
5.758GluAsn: 5.758 ± 1.843
0.64GluPro: 0.64 ± 0.627
5.118GluGln: 5.118 ± 2.234
5.118GluArg: 5.118 ± 3.139
4.479GluSer: 4.479 ± 2.279
4.479GluThr: 4.479 ± 2.2
2.559GluVal: 2.559 ± 1.801
0.64GluTrp: 0.64 ± 0.627
3.839GluTyr: 3.839 ± 1.59
0.0GluXaa: 0.0 ± 0.0
Phe
1.919PheAla: 1.919 ± 0.862
0.0PheCys: 0.0 ± 0.0
0.64PheAsp: 0.64 ± 0.416
1.919PheGlu: 1.919 ± 1.249
0.0PhePhe: 0.0 ± 0.0
2.559PheGly: 2.559 ± 1.262
0.0PheHis: 0.0 ± 0.0
1.28PheIle: 1.28 ± 1.138
3.839PheLys: 3.839 ± 1.858
1.919PheLeu: 1.919 ± 1.073
0.0PheMet: 0.0 ± 0.0
2.559PheAsn: 2.559 ± 1.381
0.64PhePro: 0.64 ± 0.416
1.919PheGln: 1.919 ± 1.112
0.0PheArg: 0.0 ± 0.0
1.28PheSer: 1.28 ± 0.631
2.559PheThr: 2.559 ± 1.665
0.0PheVal: 0.0 ± 0.0
0.64PheTrp: 0.64 ± 0.416
0.0PheTyr: 0.0 ± 0.0
0.0PheXaa: 0.0 ± 0.0
Gly
6.398GlyAla: 6.398 ± 3.366
0.0GlyCys: 0.0 ± 0.0
3.199GlyAsp: 3.199 ± 1.414
1.28GlyGlu: 1.28 ± 1.255
1.28GlyPhe: 1.28 ± 0.832
7.678GlyGly: 7.678 ± 3.391
1.919GlyHis: 1.919 ± 1.112
8.317GlyIle: 8.317 ± 2.187
6.398GlyLys: 6.398 ± 1.602
4.479GlyLeu: 4.479 ± 2.045
2.559GlyMet: 2.559 ± 1.801
3.199GlyAsn: 3.199 ± 1.054
0.64GlyPro: 0.64 ± 0.416
3.839GlyGln: 3.839 ± 2.255
0.64GlyArg: 0.64 ± 0.416
5.758GlySer: 5.758 ± 2.765
3.839GlyThr: 3.839 ± 1.52
4.479GlyVal: 4.479 ± 1.762
3.839GlyTrp: 3.839 ± 2.162
3.839GlyTyr: 3.839 ± 2.318
0.0GlyXaa: 0.0 ± 0.0
His
1.28HisAla: 1.28 ± 1.097
0.0HisCys: 0.0 ± 0.0
1.28HisAsp: 1.28 ± 1.097
1.28HisGlu: 1.28 ± 0.781
0.0HisPhe: 0.0 ± 0.0
1.28HisGly: 1.28 ± 0.559
1.919HisHis: 1.919 ± 0.945
1.28HisIle: 1.28 ± 0.832
3.199HisLys: 3.199 ± 1.645
1.28HisLeu: 1.28 ± 0.559
1.28HisMet: 1.28 ± 0.516
1.28HisAsn: 1.28 ± 1.254
1.28HisPro: 1.28 ± 0.559
0.0HisGln: 0.0 ± 0.0
0.64HisArg: 0.64 ± 1.058
1.28HisSer: 1.28 ± 0.985
1.28HisThr: 1.28 ± 0.832
0.0HisVal: 0.0 ± 0.0
1.919HisTrp: 1.919 ± 1.601
0.64HisTyr: 0.64 ± 0.627
0.0HisXaa: 0.0 ± 0.0
Ile
8.317IleAla: 8.317 ± 2.023
0.0IleCys: 0.0 ± 0.0
5.118IleAsp: 5.118 ± 2.182
7.038IleGlu: 7.038 ± 3.169
0.64IlePhe: 0.64 ± 0.416
5.118IleGly: 5.118 ± 1.504
2.559IleHis: 2.559 ± 0.915
2.559IleIle: 2.559 ± 1.577
12.156IleLys: 12.156 ± 2.789
5.758IleLeu: 5.758 ± 1.503
0.64IleMet: 0.64 ± 1.171
4.479IleAsn: 4.479 ± 1.604
3.839IlePro: 3.839 ± 1.291
1.919IleGln: 1.919 ± 0.862
6.398IleArg: 6.398 ± 2.902
3.199IleSer: 3.199 ± 1.054
7.678IleThr: 7.678 ± 2.178
2.559IleVal: 2.559 ± 0.861
1.28IleTrp: 1.28 ± 0.559
2.559IleTyr: 2.559 ± 1.117
0.0IleXaa: 0.0 ± 0.0
Lys
5.118LysAla: 5.118 ± 1.452
0.0LysCys: 0.0 ± 0.0
4.479LysAsp: 4.479 ± 1.172
8.317LysGlu: 8.317 ± 3.738
2.559LysPhe: 2.559 ± 1.037
3.199LysGly: 3.199 ± 1.414
1.28LysHis: 1.28 ± 1.254
9.597LysIle: 9.597 ± 2.661
8.317LysLys: 8.317 ± 5.781
8.957LysLeu: 8.957 ± 1.631
3.839LysMet: 3.839 ± 1.55
4.479LysAsn: 4.479 ± 1.349
1.919LysPro: 1.919 ± 0.988
3.839LysGln: 3.839 ± 1.59
3.839LysArg: 3.839 ± 2.892
3.199LysSer: 3.199 ± 0.702
7.678LysThr: 7.678 ± 2.681
1.919LysVal: 1.919 ± 1.073
0.0LysTrp: 0.0 ± 0.0
4.479LysTyr: 4.479 ± 1.805
0.0LysXaa: 0.0 ± 0.0
Leu
3.839LeuAla: 3.839 ± 1.056
0.0LeuCys: 0.0 ± 0.0
6.398LeuAsp: 6.398 ± 1.413
5.758LeuGlu: 5.758 ± 2.117
2.559LeuPhe: 2.559 ± 1.198
5.118LeuGly: 5.118 ± 1.334
4.479LeuHis: 4.479 ± 1.805
3.199LeuIle: 3.199 ± 1.873
4.479LeuLys: 4.479 ± 4.278
7.038LeuLeu: 7.038 ± 1.818
1.28LeuMet: 1.28 ± 0.631
5.118LeuAsn: 5.118 ± 0.697
1.28LeuPro: 1.28 ± 0.832
3.199LeuGln: 3.199 ± 1.163
1.919LeuArg: 1.919 ± 1.267
1.919LeuSer: 1.919 ± 0.595
8.317LeuThr: 8.317 ± 1.785
2.559LeuVal: 2.559 ± 0.861
1.28LeuTrp: 1.28 ± 0.781
3.199LeuTyr: 3.199 ± 1.667
0.0LeuXaa: 0.0 ± 0.0
Met
1.919MetAla: 1.919 ± 0.988
0.0MetCys: 0.0 ± 0.0
2.559MetAsp: 2.559 ± 0.915
1.919MetGlu: 1.919 ± 0.595
0.0MetPhe: 0.0 ± 0.0
0.0MetGly: 0.0 ± 0.0
0.0MetHis: 0.0 ± 0.0
0.64MetIle: 0.64 ± 0.416
0.0MetLys: 0.0 ± 0.0
1.28MetLeu: 1.28 ± 0.631
1.919MetMet: 1.919 ± 0.822
1.28MetAsn: 1.28 ± 0.832
0.0MetPro: 0.0 ± 0.0
1.919MetGln: 1.919 ± 1.896
1.28MetArg: 1.28 ± 0.631
4.479MetSer: 4.479 ± 1.544
1.919MetThr: 1.919 ± 1.275
2.559MetVal: 2.559 ± 1.198
0.64MetTrp: 0.64 ± 0.632
2.559MetTyr: 2.559 ± 0.99
0.0MetXaa: 0.0 ± 0.0
Asn
3.839AsnAla: 3.839 ± 1.138
0.0AsnCys: 0.0 ± 0.0
3.839AsnAsp: 3.839 ± 0.766
6.398AsnGlu: 6.398 ± 2.789
2.559AsnPhe: 2.559 ± 1.091
8.317AsnGly: 8.317 ± 2.729
0.64AsnHis: 0.64 ± 0.416
4.479AsnIle: 4.479 ± 1.015
5.758AsnLys: 5.758 ± 2.419
3.839AsnLeu: 3.839 ± 1.893
0.64AsnMet: 0.64 ± 0.555
3.839AsnAsn: 3.839 ± 1.424
1.28AsnPro: 1.28 ± 0.985
3.199AsnGln: 3.199 ± 1.153
2.559AsnArg: 2.559 ± 0.667
1.28AsnSer: 1.28 ± 0.559
5.118AsnThr: 5.118 ± 0.947
0.64AsnVal: 0.64 ± 1.058
1.919AsnTrp: 1.919 ± 1.081
3.839AsnTyr: 3.839 ± 1.056
0.0AsnXaa: 0.0 ± 0.0
Pro
1.919ProAla: 1.919 ± 1.249
0.0ProCys: 0.0 ± 0.0
0.64ProAsp: 0.64 ± 1.171
1.919ProGlu: 1.919 ± 1.073
1.28ProPhe: 1.28 ± 0.985
0.0ProGly: 0.0 ± 0.0
0.0ProHis: 0.0 ± 0.0
3.199ProIle: 3.199 ± 1.645
0.64ProLys: 0.64 ± 1.058
3.199ProLeu: 3.199 ± 2.081
0.0ProMet: 0.0 ± 0.0
1.28ProAsn: 1.28 ± 0.832
0.64ProPro: 0.64 ± 0.416
0.0ProGln: 0.0 ± 0.0
3.199ProArg: 3.199 ± 1.466
1.28ProSer: 1.28 ± 0.631
1.919ProThr: 1.919 ± 0.862
1.28ProVal: 1.28 ± 0.832
0.64ProTrp: 0.64 ± 0.416
0.64ProTyr: 0.64 ± 1.058
0.0ProXaa: 0.0 ± 0.0
Gln
3.839GlnAla: 3.839 ± 3.793
0.0GlnCys: 0.0 ± 0.0
1.28GlnAsp: 1.28 ± 0.559
4.479GlnGlu: 4.479 ± 1.034
1.28GlnPhe: 1.28 ± 0.631
1.28GlnGly: 1.28 ± 0.832
1.28GlnHis: 1.28 ± 0.559
5.118GlnIle: 5.118 ± 1.76
4.479GlnLys: 4.479 ± 2.532
4.479GlnLeu: 4.479 ± 1.78
1.919GlnMet: 1.919 ± 0.796
1.919GlnAsn: 1.919 ± 0.988
1.919GlnPro: 1.919 ± 1.249
5.118GlnGln: 5.118 ± 2.18
2.559GlnArg: 2.559 ± 1.464
1.919GlnSer: 1.919 ± 0.862
2.559GlnThr: 2.559 ± 1.341
5.118GlnVal: 5.118 ± 1.591
0.0GlnTrp: 0.0 ± 0.0
1.28GlnTyr: 1.28 ± 0.781
0.0GlnXaa: 0.0 ± 0.0
Arg
1.919ArgAla: 1.919 ± 0.595
0.0ArgCys: 0.0 ± 0.0
1.919ArgAsp: 1.919 ± 0.595
1.919ArgGlu: 1.919 ± 1.112
0.64ArgPhe: 0.64 ± 0.627
1.919ArgGly: 1.919 ± 0.595
1.919ArgHis: 1.919 ± 1.601
4.479ArgIle: 4.479 ± 1.461
5.758ArgLys: 5.758 ± 2.727
5.118ArgLeu: 5.118 ± 1.686
1.919ArgMet: 1.919 ± 0.595
1.919ArgAsn: 1.919 ± 0.945
0.64ArgPro: 0.64 ± 1.171
2.559ArgGln: 2.559 ± 2.193
2.559ArgArg: 2.559 ± 1.373
2.559ArgSer: 2.559 ± 0.99
4.479ArgThr: 4.479 ± 0.905
0.64ArgVal: 0.64 ± 0.416
1.919ArgTrp: 1.919 ± 1.267
1.28ArgTyr: 1.28 ± 0.559
0.0ArgXaa: 0.0 ± 0.0
Ser
5.758SerAla: 5.758 ± 0.873
1.28SerCys: 1.28 ± 0.559
1.28SerAsp: 1.28 ± 0.832
3.199SerGlu: 3.199 ± 2.428
1.919SerPhe: 1.919 ± 0.862
7.678SerGly: 7.678 ± 3.944
0.0SerHis: 0.0 ± 0.0
3.839SerIle: 3.839 ± 2.309
3.199SerLys: 3.199 ± 1.136
2.559SerLeu: 2.559 ± 1.047
0.0SerMet: 0.0 ± 0.0
3.199SerAsn: 3.199 ± 0.939
0.0SerPro: 0.0 ± 0.0
3.199SerGln: 3.199 ± 1.283
1.28SerArg: 1.28 ± 1.097
0.64SerSer: 0.64 ± 0.627
4.479SerThr: 4.479 ± 1.777
0.64SerVal: 0.64 ± 0.416
0.0SerTrp: 0.0 ± 0.0
0.64SerTyr: 0.64 ± 0.416
0.0SerXaa: 0.0 ± 0.0
Thr
1.919ThrAla: 1.919 ± 1.249
0.64ThrCys: 0.64 ± 0.627
2.559ThrAsp: 2.559 ± 0.667
7.678ThrGlu: 7.678 ± 2.148
1.28ThrPhe: 1.28 ± 0.631
7.038ThrGly: 7.038 ± 2.73
1.919ThrHis: 1.919 ± 0.862
6.398ThrIle: 6.398 ± 2.26
5.118ThrLys: 5.118 ± 1.934
3.839ThrLeu: 3.839 ± 1.6
1.919ThrMet: 1.919 ± 0.862
7.678ThrAsn: 7.678 ± 2.344
1.28ThrPro: 1.28 ± 0.985
4.479ThrGln: 4.479 ± 1.544
2.559ThrArg: 2.559 ± 1.117
2.559ThrSer: 2.559 ± 1.037
6.398ThrThr: 6.398 ± 2.904
2.559ThrVal: 2.559 ± 1.309
0.0ThrTrp: 0.0 ± 0.0
5.758ThrTyr: 5.758 ± 2.399
0.0ThrXaa: 0.0 ± 0.0
Val
1.28ValAla: 1.28 ± 1.264
0.0ValCys: 0.0 ± 0.0
5.758ValAsp: 5.758 ± 1.761
1.919ValGlu: 1.919 ± 0.862
3.199ValPhe: 3.199 ± 2.081
1.919ValGly: 1.919 ± 1.896
0.0ValHis: 0.0 ± 0.0
4.479ValIle: 4.479 ± 1.035
0.64ValLys: 0.64 ± 1.058
1.28ValLeu: 1.28 ± 0.985
0.64ValMet: 0.64 ± 0.416
3.839ValAsn: 3.839 ± 2.061
1.28ValPro: 1.28 ± 0.832
1.919ValGln: 1.919 ± 1.896
2.559ValArg: 2.559 ± 1.717
2.559ValSer: 2.559 ± 0.667
3.199ValThr: 3.199 ± 0.966
0.0ValVal: 0.0 ± 0.0
0.64ValTrp: 0.64 ± 0.627
0.64ValTyr: 0.64 ± 0.627
0.0ValXaa: 0.0 ± 0.0
Trp
0.64TrpAla: 0.64 ± 0.416
0.0TrpCys: 0.0 ± 0.0
4.479TrpAsp: 4.479 ± 1.528
0.64TrpGlu: 0.64 ± 0.632
0.0TrpPhe: 0.0 ± 0.0
0.64TrpGly: 0.64 ± 0.632
0.0TrpHis: 0.0 ± 0.0
1.28TrpIle: 1.28 ± 0.559
0.64TrpLys: 0.64 ± 0.632
3.839TrpLeu: 3.839 ± 1.056
0.64TrpMet: 0.64 ± 0.416
0.64TrpAsn: 0.64 ± 0.627
0.64TrpPro: 0.64 ± 0.627
0.64TrpGln: 0.64 ± 0.627
1.28TrpArg: 1.28 ± 1.254
1.28TrpSer: 1.28 ± 0.985
1.28TrpThr: 1.28 ± 0.559
1.28TrpVal: 1.28 ± 0.559
0.0TrpTrp: 0.0 ± 0.0
0.64TrpTyr: 0.64 ± 1.058
0.0TrpXaa: 0.0 ± 0.0
Tyr
1.28TyrAla: 1.28 ± 0.832
0.64TyrCys: 0.64 ± 0.416
3.839TyrAsp: 3.839 ± 2.497
3.839TyrGlu: 3.839 ± 2.34
0.64TyrPhe: 0.64 ± 0.416
3.199TyrGly: 3.199 ± 1.117
0.0TyrHis: 0.0 ± 0.0
4.479TyrIle: 4.479 ± 1.461
5.118TyrLys: 5.118 ± 2.746
0.64TyrLeu: 0.64 ± 0.416
1.28TyrMet: 1.28 ± 0.559
4.479TyrAsn: 4.479 ± 1.845
1.28TyrPro: 1.28 ± 0.559
1.919TyrGln: 1.919 ± 0.862
1.28TyrArg: 1.28 ± 0.559
2.559TyrSer: 2.559 ± 1.381
0.0TyrThr: 0.0 ± 0.0
2.559TyrVal: 2.559 ± 1.717
1.28TyrTrp: 1.28 ± 0.559
2.559TyrTyr: 2.559 ± 1.117
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 5 proteins (1564 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski