Amino acid dipepetide frequency for Tortoise microvirus 78

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
11.673AlaAla: 11.673 ± 3.372
0.0AlaCys: 0.0 ± 0.0
2.779AlaAsp: 2.779 ± 1.194
5.003AlaGlu: 5.003 ± 2.186
5.003AlaPhe: 5.003 ± 0.985
11.673AlaGly: 11.673 ± 2.667
2.779AlaHis: 2.779 ± 1.104
4.447AlaIle: 4.447 ± 1.325
7.782AlaLys: 7.782 ± 2.293
6.67AlaLeu: 6.67 ± 1.602
2.223AlaMet: 2.223 ± 0.574
3.335AlaAsn: 3.335 ± 0.999
10.006AlaPro: 10.006 ± 1.536
6.115AlaGln: 6.115 ± 2.213
6.67AlaArg: 6.67 ± 3.737
3.891AlaSer: 3.891 ± 1.175
5.559AlaThr: 5.559 ± 1.535
7.782AlaVal: 7.782 ± 1.915
1.112AlaTrp: 1.112 ± 0.612
3.335AlaTyr: 3.335 ± 0.865
0.0AlaXaa: 0.0 ± 0.0
Cys
0.0CysAla: 0.0 ± 0.0
0.0CysCys: 0.0 ± 0.0
0.0CysAsp: 0.0 ± 0.0
0.0CysGlu: 0.0 ± 0.0
0.0CysPhe: 0.0 ± 0.0
0.0CysGly: 0.0 ± 0.0
0.0CysHis: 0.0 ± 0.0
0.0CysIle: 0.0 ± 0.0
0.556CysLys: 0.556 ± 0.636
0.0CysLeu: 0.0 ± 0.0
0.0CysMet: 0.0 ± 0.0
0.0CysAsn: 0.0 ± 0.0
0.0CysPro: 0.0 ± 0.0
0.0CysGln: 0.0 ± 0.0
0.0CysArg: 0.0 ± 0.0
0.556CysSer: 0.556 ± 0.636
0.0CysThr: 0.0 ± 0.0
0.0CysVal: 0.0 ± 0.0
0.0CysTrp: 0.0 ± 0.0
0.0CysTyr: 0.0 ± 0.0
0.0CysXaa: 0.0 ± 0.0
Asp
5.559AspAla: 5.559 ± 1.44
0.0AspCys: 0.0 ± 0.0
5.559AspAsp: 5.559 ± 2.283
1.668AspGlu: 1.668 ± 1.211
1.668AspPhe: 1.668 ± 0.914
6.115AspGly: 6.115 ± 1.23
0.556AspHis: 0.556 ± 0.633
1.112AspIle: 1.112 ± 0.531
1.668AspLys: 1.668 ± 0.944
3.335AspLeu: 3.335 ± 1.34
0.556AspMet: 0.556 ± 0.404
0.0AspAsn: 0.0 ± 0.0
7.226AspPro: 7.226 ± 1.047
3.335AspGln: 3.335 ± 1.456
3.891AspArg: 3.891 ± 0.847
2.223AspSer: 2.223 ± 0.921
3.335AspThr: 3.335 ± 1.143
5.003AspVal: 5.003 ± 1.269
2.223AspTrp: 2.223 ± 1.074
2.223AspTyr: 2.223 ± 0.57
0.0AspXaa: 0.0 ± 0.0
Glu
5.003GluAla: 5.003 ± 1.717
0.0GluCys: 0.0 ± 0.0
1.668GluAsp: 1.668 ± 0.732
2.223GluGlu: 2.223 ± 1.088
1.668GluPhe: 1.668 ± 0.621
5.003GluGly: 5.003 ± 1.157
0.556GluHis: 0.556 ± 0.493
1.112GluIle: 1.112 ± 0.755
2.223GluLys: 2.223 ± 0.65
8.338GluLeu: 8.338 ± 1.895
2.223GluMet: 2.223 ± 0.807
1.112GluAsn: 1.112 ± 0.807
2.779GluPro: 2.779 ± 1.567
3.335GluGln: 3.335 ± 1.297
6.115GluArg: 6.115 ± 1.097
3.891GluSer: 3.891 ± 1.13
2.779GluThr: 2.779 ± 1.308
7.782GluVal: 7.782 ± 1.701
1.112GluTrp: 1.112 ± 0.531
1.668GluTyr: 1.668 ± 1.48
0.0GluXaa: 0.0 ± 0.0
Phe
2.223PheAla: 2.223 ± 0.711
0.0PheCys: 0.0 ± 0.0
3.335PheAsp: 3.335 ± 0.966
2.779PheGlu: 2.779 ± 0.674
0.556PhePhe: 0.556 ± 0.404
3.891PheGly: 3.891 ± 1.62
0.556PheHis: 0.556 ± 0.404
1.668PheIle: 1.668 ± 0.921
0.556PheLys: 0.556 ± 0.493
1.668PheLeu: 1.668 ± 0.596
0.556PheMet: 0.556 ± 0.404
1.668PheAsn: 1.668 ± 0.732
1.668PhePro: 1.668 ± 0.708
1.112PheGln: 1.112 ± 0.473
1.112PheArg: 1.112 ± 0.531
0.556PheSer: 0.556 ± 0.493
1.112PheThr: 1.112 ± 0.958
2.223PheVal: 2.223 ± 0.638
1.668PheTrp: 1.668 ± 0.942
2.223PheTyr: 2.223 ± 0.984
0.0PheXaa: 0.0 ± 0.0
Gly
8.338GlyAla: 8.338 ± 1.983
0.0GlyCys: 0.0 ± 0.0
5.003GlyAsp: 5.003 ± 2.709
7.226GlyGlu: 7.226 ± 3.133
2.779GlyPhe: 2.779 ± 0.555
15.008GlyGly: 15.008 ± 3.336
1.112GlyHis: 1.112 ± 0.722
5.003GlyIle: 5.003 ± 0.986
3.335GlyLys: 3.335 ± 1.301
7.226GlyLeu: 7.226 ± 2.211
2.223GlyMet: 2.223 ± 0.894
5.003GlyAsn: 5.003 ± 1.733
6.67GlyPro: 6.67 ± 2.802
3.891GlyGln: 3.891 ± 1.236
5.003GlyArg: 5.003 ± 1.663
5.559GlySer: 5.559 ± 1.768
9.45GlyThr: 9.45 ± 2.381
6.115GlyVal: 6.115 ± 1.263
2.223GlyTrp: 2.223 ± 1.571
2.779GlyTyr: 2.779 ± 0.859
0.0GlyXaa: 0.0 ± 0.0
His
1.668HisAla: 1.668 ± 0.936
0.0HisCys: 0.0 ± 0.0
0.0HisAsp: 0.0 ± 0.0
1.668HisGlu: 1.668 ± 0.803
0.0HisPhe: 0.0 ± 0.0
2.223HisGly: 2.223 ± 0.717
0.0HisHis: 0.0 ± 0.0
0.556HisIle: 0.556 ± 0.404
1.112HisLys: 1.112 ± 0.613
2.779HisLeu: 2.779 ± 1.3
0.0HisMet: 0.0 ± 0.0
0.556HisAsn: 0.556 ± 0.404
0.556HisPro: 0.556 ± 0.647
0.556HisGln: 0.556 ± 0.647
0.0HisArg: 0.0 ± 0.0
1.112HisSer: 1.112 ± 1.271
0.0HisThr: 0.0 ± 0.0
2.223HisVal: 2.223 ± 0.928
2.779HisTrp: 2.779 ± 1.331
0.556HisTyr: 0.556 ± 0.493
0.0HisXaa: 0.0 ± 0.0
Ile
1.668IleAla: 1.668 ± 1.262
0.0IleCys: 0.0 ± 0.0
1.668IleAsp: 1.668 ± 1.211
2.223IleGlu: 2.223 ± 0.726
0.0IlePhe: 0.0 ± 0.0
3.335IleGly: 3.335 ± 1.299
1.112IleHis: 1.112 ± 0.531
0.556IleIle: 0.556 ± 0.487
2.223IleLys: 2.223 ± 0.93
3.891IleLeu: 3.891 ± 1.046
1.112IleMet: 1.112 ± 0.613
0.556IleAsn: 0.556 ± 0.633
3.335IlePro: 3.335 ± 0.756
0.0IleGln: 0.0 ± 0.0
2.223IleArg: 2.223 ± 1.407
0.556IleSer: 0.556 ± 0.479
1.668IleThr: 1.668 ± 1.06
4.447IleVal: 4.447 ± 2.105
1.112IleTrp: 1.112 ± 0.807
0.0IleTyr: 0.0 ± 0.0
0.0IleXaa: 0.0 ± 0.0
Lys
6.115LysAla: 6.115 ± 2.322
0.0LysCys: 0.0 ± 0.0
2.779LysAsp: 2.779 ± 1.311
2.779LysGlu: 2.779 ± 1.119
1.668LysPhe: 1.668 ± 0.596
3.335LysGly: 3.335 ± 1.304
0.556LysHis: 0.556 ± 0.404
1.668LysIle: 1.668 ± 0.732
3.335LysLys: 3.335 ± 1.119
3.891LysLeu: 3.891 ± 2.096
1.668LysMet: 1.668 ± 1.27
0.556LysAsn: 0.556 ± 0.404
3.335LysPro: 3.335 ± 0.751
2.223LysGln: 2.223 ± 0.921
4.447LysArg: 4.447 ± 1.274
0.0LysSer: 0.0 ± 0.0
1.668LysThr: 1.668 ± 1.211
3.335LysVal: 3.335 ± 0.856
0.556LysTrp: 0.556 ± 0.404
1.112LysTyr: 1.112 ± 0.722
0.0LysXaa: 0.0 ± 0.0
Leu
12.229LeuAla: 12.229 ± 2.001
0.0LeuCys: 0.0 ± 0.0
6.115LeuAsp: 6.115 ± 1.105
5.003LeuGlu: 5.003 ± 0.854
1.112LeuPhe: 1.112 ± 0.69
9.45LeuGly: 9.45 ± 2.274
2.223LeuHis: 2.223 ± 1.127
1.112LeuIle: 1.112 ± 1.267
3.891LeuLys: 3.891 ± 0.933
2.779LeuLeu: 2.779 ± 1.047
2.779LeuMet: 2.779 ± 1.047
4.447LeuAsn: 4.447 ± 1.299
2.779LeuPro: 2.779 ± 0.713
2.223LeuGln: 2.223 ± 1.407
7.226LeuArg: 7.226 ± 2.059
6.115LeuSer: 6.115 ± 1.893
1.668LeuThr: 1.668 ± 0.818
3.891LeuVal: 3.891 ± 1.159
0.556LeuTrp: 0.556 ± 0.487
0.0LeuTyr: 0.0 ± 0.0
0.0LeuXaa: 0.0 ± 0.0
Met
3.891MetAla: 3.891 ± 1.579
0.0MetCys: 0.0 ± 0.0
1.668MetAsp: 1.668 ± 0.665
1.112MetGlu: 1.112 ± 0.672
1.112MetPhe: 1.112 ± 0.772
3.335MetGly: 3.335 ± 1.369
0.556MetHis: 0.556 ± 0.404
0.556MetIle: 0.556 ± 0.647
0.556MetLys: 0.556 ± 0.404
1.668MetLeu: 1.668 ± 0.708
0.0MetMet: 0.0 ± 0.0
1.668MetAsn: 1.668 ± 0.811
1.668MetPro: 1.668 ± 0.639
0.556MetGln: 0.556 ± 0.647
2.223MetArg: 2.223 ± 1.022
0.556MetSer: 0.556 ± 0.493
1.112MetThr: 1.112 ± 0.755
0.556MetVal: 0.556 ± 0.404
0.0MetTrp: 0.0 ± 0.0
0.556MetTyr: 0.556 ± 0.493
0.0MetXaa: 0.0 ± 0.0
Asn
1.112AsnAla: 1.112 ± 0.473
0.0AsnCys: 0.0 ± 0.0
1.668AsnAsp: 1.668 ± 0.99
1.668AsnGlu: 1.668 ± 0.862
1.668AsnPhe: 1.668 ± 1.211
5.559AsnGly: 5.559 ± 2.015
1.668AsnHis: 1.668 ± 1.211
1.112AsnIle: 1.112 ± 0.69
1.668AsnLys: 1.668 ± 0.921
3.335AsnLeu: 3.335 ± 0.998
0.556AsnMet: 0.556 ± 0.479
0.556AsnAsn: 0.556 ± 0.636
5.003AsnPro: 5.003 ± 1.294
1.112AsnGln: 1.112 ± 0.53
3.891AsnArg: 3.891 ± 1.243
3.335AsnSer: 3.335 ± 1.261
2.223AsnThr: 2.223 ± 1.248
3.891AsnVal: 3.891 ± 1.365
0.556AsnTrp: 0.556 ± 0.647
0.0AsnTyr: 0.0 ± 0.0
0.0AsnXaa: 0.0 ± 0.0
Pro
6.115ProAla: 6.115 ± 1.391
0.0ProCys: 0.0 ± 0.0
3.891ProAsp: 3.891 ± 1.669
6.115ProGlu: 6.115 ± 2.146
2.223ProPhe: 2.223 ± 1.615
9.45ProGly: 9.45 ± 4.169
1.112ProHis: 1.112 ± 0.722
1.668ProIle: 1.668 ± 0.639
1.668ProLys: 1.668 ± 0.944
5.559ProLeu: 5.559 ± 1.241
2.223ProMet: 2.223 ± 0.599
2.223ProAsn: 2.223 ± 1.156
2.779ProPro: 2.779 ± 1.198
4.447ProGln: 4.447 ± 1.022
4.447ProArg: 4.447 ± 0.753
4.447ProSer: 4.447 ± 1.816
3.335ProThr: 3.335 ± 1.264
5.559ProVal: 5.559 ± 2.048
1.668ProTrp: 1.668 ± 0.717
0.556ProTyr: 0.556 ± 0.487
0.0ProXaa: 0.0 ± 0.0
Gln
7.782GlnAla: 7.782 ± 1.686
0.0GlnCys: 0.0 ± 0.0
1.112GlnAsp: 1.112 ± 0.531
2.779GlnGlu: 2.779 ± 0.961
0.556GlnPhe: 0.556 ± 0.404
2.779GlnGly: 2.779 ± 1.314
1.112GlnHis: 1.112 ± 0.531
0.556GlnIle: 0.556 ± 0.404
3.335GlnLys: 3.335 ± 1.297
2.779GlnLeu: 2.779 ± 0.707
0.0GlnMet: 0.0 ± 0.0
3.891GlnAsn: 3.891 ± 1.123
5.003GlnPro: 5.003 ± 1.77
2.223GlnGln: 2.223 ± 0.784
2.223GlnArg: 2.223 ± 1.167
3.335GlnSer: 3.335 ± 1.011
2.223GlnThr: 2.223 ± 0.65
3.891GlnVal: 3.891 ± 1.892
0.0GlnTrp: 0.0 ± 0.0
2.223GlnTyr: 2.223 ± 0.57
0.0GlnXaa: 0.0 ± 0.0
Arg
11.117ArgAla: 11.117 ± 2.249
0.0ArgCys: 0.0 ± 0.0
3.335ArgAsp: 3.335 ± 1.053
4.447ArgGlu: 4.447 ± 1.458
3.335ArgPhe: 3.335 ± 1.088
3.891ArgGly: 3.891 ± 1.384
0.556ArgHis: 0.556 ± 0.493
2.779ArgIle: 2.779 ± 1.533
1.112ArgLys: 1.112 ± 0.934
6.67ArgLeu: 6.67 ± 0.893
1.112ArgMet: 1.112 ± 0.751
2.223ArgAsn: 2.223 ± 0.921
3.891ArgPro: 3.891 ± 1.222
6.115ArgGln: 6.115 ± 1.015
4.447ArgArg: 4.447 ± 1.982
2.223ArgSer: 2.223 ± 0.717
3.335ArgThr: 3.335 ± 1.735
1.112ArgVal: 1.112 ± 0.889
1.668ArgTrp: 1.668 ± 0.798
1.668ArgTyr: 1.668 ± 0.942
0.0ArgXaa: 0.0 ± 0.0
Ser
6.115SerAla: 6.115 ± 2.748
0.0SerCys: 0.0 ± 0.0
5.003SerAsp: 5.003 ± 1.706
3.335SerGlu: 3.335 ± 1.265
2.223SerPhe: 2.223 ± 1.088
5.003SerGly: 5.003 ± 1.307
1.112SerHis: 1.112 ± 0.68
2.223SerIle: 2.223 ± 1.38
2.223SerLys: 2.223 ± 1.147
2.223SerLeu: 2.223 ± 1.376
1.668SerMet: 1.668 ± 0.753
2.223SerAsn: 2.223 ± 0.785
1.112SerPro: 1.112 ± 0.613
2.779SerGln: 2.779 ± 0.928
1.668SerArg: 1.668 ± 0.596
2.779SerSer: 2.779 ± 1.306
1.112SerThr: 1.112 ± 0.726
1.668SerVal: 1.668 ± 1.211
2.223SerTrp: 2.223 ± 1.375
2.223SerTyr: 2.223 ± 0.815
0.0SerXaa: 0.0 ± 0.0
Thr
6.67ThrAla: 6.67 ± 2.135
0.0ThrCys: 0.0 ± 0.0
5.559ThrAsp: 5.559 ± 2.318
2.779ThrGlu: 2.779 ± 1.276
1.668ThrPhe: 1.668 ± 0.803
2.779ThrGly: 2.779 ± 1.091
0.556ThrHis: 0.556 ± 0.594
0.0ThrIle: 0.0 ± 0.0
0.556ThrLys: 0.556 ± 0.493
5.003ThrLeu: 5.003 ± 0.989
0.0ThrMet: 0.0 ± 0.0
1.668ThrAsn: 1.668 ± 1.06
3.891ThrPro: 3.891 ± 1.28
3.335ThrGln: 3.335 ± 1.134
1.668ThrArg: 1.668 ± 0.69
1.668ThrSer: 1.668 ± 0.872
3.335ThrThr: 3.335 ± 1.464
3.891ThrVal: 3.891 ± 1.877
3.335ThrTrp: 3.335 ± 1.038
1.112ThrTyr: 1.112 ± 0.987
0.0ThrXaa: 0.0 ± 0.0
Val
5.559ValAla: 5.559 ± 1.33
0.556ValCys: 0.556 ± 0.636
3.335ValAsp: 3.335 ± 1.052
6.115ValGlu: 6.115 ± 2.098
1.112ValPhe: 1.112 ± 0.53
6.115ValGly: 6.115 ± 1.182
1.668ValHis: 1.668 ± 0.861
5.003ValIle: 5.003 ± 1.526
2.779ValLys: 2.779 ± 0.878
3.891ValLeu: 3.891 ± 1.003
2.779ValMet: 2.779 ± 0.989
5.003ValAsn: 5.003 ± 1.378
5.559ValPro: 5.559 ± 2.413
3.335ValGln: 3.335 ± 1.376
3.891ValArg: 3.891 ± 1.375
3.891ValSer: 3.891 ± 1.346
3.335ValThr: 3.335 ± 1.381
7.226ValVal: 7.226 ± 2.453
1.112ValTrp: 1.112 ± 0.473
1.112ValTyr: 1.112 ± 0.531
0.0ValXaa: 0.0 ± 0.0
Trp
2.779TrpAla: 2.779 ± 1.447
0.556TrpCys: 0.556 ± 0.636
0.556TrpAsp: 0.556 ± 0.493
1.112TrpGlu: 1.112 ± 0.722
1.112TrpPhe: 1.112 ± 0.473
2.223TrpGly: 2.223 ± 1.64
0.556TrpHis: 0.556 ± 0.493
0.0TrpIle: 0.0 ± 0.0
2.779TrpLys: 2.779 ± 1.91
1.668TrpLeu: 1.668 ± 1.262
0.556TrpMet: 0.556 ± 0.468
1.668TrpAsn: 1.668 ± 0.936
1.668TrpPro: 1.668 ± 0.942
0.556TrpGln: 0.556 ± 0.493
2.779TrpArg: 2.779 ± 1.291
1.112TrpSer: 1.112 ± 0.975
1.112TrpThr: 1.112 ± 0.473
1.112TrpVal: 1.112 ± 0.68
0.556TrpTrp: 0.556 ± 0.493
0.556TrpTyr: 0.556 ± 0.487
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.223TyrAla: 2.223 ± 1.061
0.0TyrCys: 0.0 ± 0.0
2.223TyrAsp: 2.223 ± 0.711
0.556TyrGlu: 0.556 ± 0.487
1.668TyrPhe: 1.668 ± 1.084
2.779TyrGly: 2.779 ± 1.562
0.0TyrHis: 0.0 ± 0.0
0.556TyrIle: 0.556 ± 0.493
1.668TyrLys: 1.668 ± 0.572
2.223TyrLeu: 2.223 ± 0.857
0.556TyrMet: 0.556 ± 0.404
2.223TyrAsn: 2.223 ± 0.946
0.556TyrPro: 0.556 ± 0.493
0.556TyrGln: 0.556 ± 0.487
1.112TyrArg: 1.112 ± 0.82
1.668TyrSer: 1.668 ± 1.117
1.112TyrThr: 1.112 ± 0.987
1.668TyrVal: 1.668 ± 0.942
0.556TyrTrp: 0.556 ± 0.493
0.556TyrTyr: 0.556 ± 0.493
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 8 proteins (1800 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski