Amino acid dipepetide frequency for Gordonia phage Bradissa

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
16.37AlaAla: 16.37 ± 1.71
1.135AlaCys: 1.135 ± 0.273
8.304AlaAsp: 8.304 ± 0.584
8.304AlaGlu: 8.304 ± 0.69
3.047AlaPhe: 3.047 ± 0.592
10.276AlaGly: 10.276 ± 0.836
2.509AlaHis: 2.509 ± 0.452
5.138AlaIle: 5.138 ± 0.546
4.541AlaLys: 4.541 ± 0.579
8.663AlaLeu: 8.663 ± 0.693
3.824AlaMet: 3.824 ± 0.709
3.525AlaAsn: 3.525 ± 0.764
5.019AlaPro: 5.019 ± 0.552
4.66AlaGln: 4.66 ± 0.576
8.245AlaArg: 8.245 ± 0.96
6.512AlaSer: 6.512 ± 0.718
7.767AlaThr: 7.767 ± 0.712
7.169AlaVal: 7.169 ± 0.658
2.211AlaTrp: 2.211 ± 0.305
2.509AlaTyr: 2.509 ± 0.392
0.0AlaXaa: 0.0 ± 0.0
Cys
0.896CysAla: 0.896 ± 0.255
0.06CysCys: 0.06 ± 0.061
0.956CysAsp: 0.956 ± 0.253
0.777CysGlu: 0.777 ± 0.265
0.06CysPhe: 0.06 ± 0.065
1.673CysGly: 1.673 ± 0.392
0.358CysHis: 0.358 ± 0.153
0.179CysIle: 0.179 ± 0.102
0.239CysLys: 0.239 ± 0.127
0.179CysLeu: 0.179 ± 0.109
0.119CysMet: 0.119 ± 0.088
0.119CysAsn: 0.119 ± 0.082
0.299CysPro: 0.299 ± 0.125
0.478CysGln: 0.478 ± 0.183
1.016CysArg: 1.016 ± 0.303
0.538CysSer: 0.538 ± 0.187
0.358CysThr: 0.358 ± 0.139
0.478CysVal: 0.478 ± 0.172
0.299CysTrp: 0.299 ± 0.168
0.358CysTyr: 0.358 ± 0.135
0.0CysXaa: 0.0 ± 0.0
Asp
7.946AspAla: 7.946 ± 0.683
0.597AspCys: 0.597 ± 0.197
5.257AspAsp: 5.257 ± 0.674
4.122AspGlu: 4.122 ± 0.55
1.912AspPhe: 1.912 ± 0.289
7.229AspGly: 7.229 ± 0.638
1.792AspHis: 1.792 ± 0.269
3.286AspIle: 3.286 ± 0.396
1.255AspLys: 1.255 ± 0.297
6.93AspLeu: 6.93 ± 0.599
1.135AspMet: 1.135 ± 0.282
1.792AspAsn: 1.792 ± 0.402
4.899AspPro: 4.899 ± 0.603
2.39AspGln: 2.39 ± 0.416
5.676AspArg: 5.676 ± 0.585
2.27AspSer: 2.27 ± 0.389
2.987AspThr: 2.987 ± 0.385
4.122AspVal: 4.122 ± 0.482
1.374AspTrp: 1.374 ± 0.292
1.733AspTyr: 1.733 ± 0.4
0.0AspXaa: 0.0 ± 0.0
Glu
6.094GluAla: 6.094 ± 0.539
0.478GluCys: 0.478 ± 0.222
3.047GluAsp: 3.047 ± 0.426
2.868GluGlu: 2.868 ± 0.409
2.211GluPhe: 2.211 ± 0.42
3.166GluGly: 3.166 ± 0.334
1.613GluHis: 1.613 ± 0.332
2.927GluIle: 2.927 ± 0.525
2.27GluLys: 2.27 ± 0.395
6.094GluLeu: 6.094 ± 0.636
1.255GluMet: 1.255 ± 0.273
1.613GluAsn: 1.613 ± 0.3
2.45GluPro: 2.45 ± 0.365
2.031GluGln: 2.031 ± 0.391
3.943GluArg: 3.943 ± 0.482
3.644GluSer: 3.644 ± 0.512
2.808GluThr: 2.808 ± 0.411
5.257GluVal: 5.257 ± 0.709
1.314GluTrp: 1.314 ± 0.324
1.733GluTyr: 1.733 ± 0.326
0.0GluXaa: 0.0 ± 0.0
Phe
2.987PheAla: 2.987 ± 0.425
0.299PheCys: 0.299 ± 0.135
2.151PheAsp: 2.151 ± 0.34
1.673PheGlu: 1.673 ± 0.367
0.418PhePhe: 0.418 ± 0.191
2.927PheGly: 2.927 ± 0.519
0.478PheHis: 0.478 ± 0.137
1.553PheIle: 1.553 ± 0.252
0.597PheLys: 0.597 ± 0.129
2.151PheLeu: 2.151 ± 0.377
0.299PheMet: 0.299 ± 0.166
0.657PheAsn: 0.657 ± 0.165
1.374PhePro: 1.374 ± 0.35
0.836PheGln: 0.836 ± 0.219
1.673PheArg: 1.673 ± 0.24
1.792PheSer: 1.792 ± 0.309
1.912PheThr: 1.912 ± 0.332
2.27PheVal: 2.27 ± 0.374
0.478PheTrp: 0.478 ± 0.143
0.657PheTyr: 0.657 ± 0.153
0.0PheXaa: 0.0 ± 0.0
Gly
9.559GlyAla: 9.559 ± 1.117
0.538GlyCys: 0.538 ± 0.221
5.616GlyAsp: 5.616 ± 0.653
4.78GlyGlu: 4.78 ± 0.501
2.39GlyPhe: 2.39 ± 0.376
9.44GlyGly: 9.44 ± 1.351
2.151GlyHis: 2.151 ± 0.401
3.644GlyIle: 3.644 ± 0.534
3.166GlyLys: 3.166 ± 0.464
5.735GlyLeu: 5.735 ± 0.686
1.374GlyMet: 1.374 ± 0.231
3.107GlyAsn: 3.107 ± 0.498
4.481GlyPro: 4.481 ± 0.539
3.585GlyGln: 3.585 ± 0.449
6.811GlyArg: 6.811 ± 0.702
4.361GlySer: 4.361 ± 0.62
6.094GlyThr: 6.094 ± 0.676
5.855GlyVal: 5.855 ± 0.563
1.912GlyTrp: 1.912 ± 0.333
1.673GlyTyr: 1.673 ± 0.272
0.0GlyXaa: 0.0 ± 0.0
His
2.091HisAla: 2.091 ± 0.391
0.597HisCys: 0.597 ± 0.192
1.135HisAsp: 1.135 ± 0.297
0.956HisGlu: 0.956 ± 0.22
0.119HisPhe: 0.119 ± 0.08
1.972HisGly: 1.972 ± 0.404
0.896HisHis: 0.896 ± 0.223
0.836HisIle: 0.836 ± 0.21
0.299HisLys: 0.299 ± 0.122
2.031HisLeu: 2.031 ± 0.401
0.179HisMet: 0.179 ± 0.109
0.777HisAsn: 0.777 ± 0.217
1.673HisPro: 1.673 ± 0.38
0.836HisGln: 0.836 ± 0.298
1.852HisArg: 1.852 ± 0.346
0.777HisSer: 0.777 ± 0.292
1.792HisThr: 1.792 ± 0.275
1.434HisVal: 1.434 ± 0.304
0.478HisTrp: 0.478 ± 0.164
0.896HisTyr: 0.896 ± 0.235
0.0HisXaa: 0.0 ± 0.0
Ile
6.393IleAla: 6.393 ± 0.587
0.06IleCys: 0.06 ± 0.058
3.525IleAsp: 3.525 ± 0.477
3.465IleGlu: 3.465 ± 0.46
0.717IlePhe: 0.717 ± 0.196
3.166IleGly: 3.166 ± 0.415
0.956IleHis: 0.956 ± 0.254
1.613IleIle: 1.613 ± 0.317
1.016IleLys: 1.016 ± 0.311
3.226IleLeu: 3.226 ± 0.445
0.358IleMet: 0.358 ± 0.122
1.314IleAsn: 1.314 ± 0.305
3.047IlePro: 3.047 ± 0.388
1.613IleGln: 1.613 ± 0.34
3.465IleArg: 3.465 ± 0.446
2.211IleSer: 2.211 ± 0.321
3.644IleThr: 3.644 ± 0.534
2.927IleVal: 2.927 ± 0.479
0.538IleTrp: 0.538 ± 0.172
1.075IleTyr: 1.075 ± 0.285
0.0IleXaa: 0.0 ± 0.0
Lys
3.465LysAla: 3.465 ± 0.534
0.299LysCys: 0.299 ± 0.14
1.733LysAsp: 1.733 ± 0.303
1.255LysGlu: 1.255 ± 0.241
0.956LysPhe: 0.956 ± 0.234
1.852LysGly: 1.852 ± 0.359
0.358LysHis: 0.358 ± 0.139
1.374LysIle: 1.374 ± 0.296
1.255LysLys: 1.255 ± 0.361
3.107LysLeu: 3.107 ± 0.384
0.538LysMet: 0.538 ± 0.182
1.374LysAsn: 1.374 ± 0.324
2.211LysPro: 2.211 ± 0.513
0.777LysGln: 0.777 ± 0.249
2.987LysArg: 2.987 ± 0.402
2.33LysSer: 2.33 ± 0.333
2.569LysThr: 2.569 ± 0.367
2.39LysVal: 2.39 ± 0.349
0.538LysTrp: 0.538 ± 0.175
0.538LysTyr: 0.538 ± 0.178
0.0LysXaa: 0.0 ± 0.0
Leu
10.216LeuAla: 10.216 ± 0.705
0.657LeuCys: 0.657 ± 0.192
6.572LeuAsp: 6.572 ± 0.71
4.66LeuGlu: 4.66 ± 0.61
2.45LeuPhe: 2.45 ± 0.402
6.154LeuGly: 6.154 ± 0.666
1.314LeuHis: 1.314 ± 0.288
3.525LeuIle: 3.525 ± 0.411
2.27LeuLys: 2.27 ± 0.424
4.361LeuLeu: 4.361 ± 0.485
1.553LeuMet: 1.553 ± 0.327
2.27LeuAsn: 2.27 ± 0.433
5.198LeuPro: 5.198 ± 0.592
2.509LeuGln: 2.509 ± 0.431
6.034LeuArg: 6.034 ± 0.872
4.302LeuSer: 4.302 ± 0.506
6.691LeuThr: 6.691 ± 0.666
4.242LeuVal: 4.242 ± 0.712
1.613LeuTrp: 1.613 ± 0.412
1.255LeuTyr: 1.255 ± 0.264
0.0LeuXaa: 0.0 ± 0.0
Met
1.912MetAla: 1.912 ± 0.315
0.418MetCys: 0.418 ± 0.175
0.896MetAsp: 0.896 ± 0.205
1.016MetGlu: 1.016 ± 0.241
0.418MetPhe: 0.418 ± 0.149
1.374MetGly: 1.374 ± 0.372
0.418MetHis: 0.418 ± 0.128
0.358MetIle: 0.358 ± 0.153
0.836MetLys: 0.836 ± 0.205
1.613MetLeu: 1.613 ± 0.275
0.418MetMet: 0.418 ± 0.162
0.657MetAsn: 0.657 ± 0.181
0.896MetPro: 0.896 ± 0.228
0.896MetGln: 0.896 ± 0.402
1.852MetArg: 1.852 ± 0.317
1.792MetSer: 1.792 ± 0.351
3.047MetThr: 3.047 ± 0.376
1.016MetVal: 1.016 ± 0.221
0.597MetTrp: 0.597 ± 0.188
0.119MetTyr: 0.119 ± 0.081
0.0MetXaa: 0.0 ± 0.0
Asn
3.824AsnAla: 3.824 ± 0.816
0.119AsnCys: 0.119 ± 0.085
1.852AsnAsp: 1.852 ± 0.309
1.255AsnGlu: 1.255 ± 0.25
1.016AsnPhe: 1.016 ± 0.29
3.764AsnGly: 3.764 ± 0.563
0.478AsnHis: 0.478 ± 0.164
1.075AsnIle: 1.075 ± 0.264
1.075AsnLys: 1.075 ± 0.212
2.33AsnLeu: 2.33 ± 0.44
0.358AsnMet: 0.358 ± 0.126
0.418AsnAsn: 0.418 ± 0.229
2.33AsnPro: 2.33 ± 0.357
1.553AsnGln: 1.553 ± 0.319
1.912AsnArg: 1.912 ± 0.37
1.792AsnSer: 1.792 ± 0.34
1.972AsnThr: 1.972 ± 0.343
1.733AsnVal: 1.733 ± 0.259
0.538AsnTrp: 0.538 ± 0.176
0.478AsnTyr: 0.478 ± 0.158
0.0AsnXaa: 0.0 ± 0.0
Pro
6.213ProAla: 6.213 ± 0.654
0.777ProCys: 0.777 ± 0.251
5.019ProAsp: 5.019 ± 0.586
4.003ProGlu: 4.003 ± 0.55
1.075ProPhe: 1.075 ± 0.209
5.377ProGly: 5.377 ± 0.564
1.494ProHis: 1.494 ± 0.431
2.211ProIle: 2.211 ± 0.389
2.091ProLys: 2.091 ± 0.35
4.003ProLeu: 4.003 ± 0.509
0.956ProMet: 0.956 ± 0.231
1.912ProAsn: 1.912 ± 0.322
3.525ProPro: 3.525 ± 0.545
1.494ProGln: 1.494 ± 0.286
4.361ProArg: 4.361 ± 0.647
2.987ProSer: 2.987 ± 0.389
3.286ProThr: 3.286 ± 0.458
3.525ProVal: 3.525 ± 0.496
1.494ProTrp: 1.494 ± 0.366
1.075ProTyr: 1.075 ± 0.228
0.0ProXaa: 0.0 ± 0.0
Gln
4.003GlnAla: 4.003 ± 0.708
0.119GlnCys: 0.119 ± 0.081
1.912GlnAsp: 1.912 ± 0.279
1.075GlnGlu: 1.075 ± 0.241
1.553GlnPhe: 1.553 ± 0.355
2.569GlnGly: 2.569 ± 0.5
1.135GlnHis: 1.135 ± 0.247
2.211GlnIle: 2.211 ± 0.377
1.016GlnLys: 1.016 ± 0.321
3.226GlnLeu: 3.226 ± 0.552
0.777GlnMet: 0.777 ± 0.2
1.016GlnAsn: 1.016 ± 0.288
1.733GlnPro: 1.733 ± 0.273
2.33GlnGln: 2.33 ± 0.703
3.166GlnArg: 3.166 ± 0.503
2.27GlnSer: 2.27 ± 0.455
2.629GlnThr: 2.629 ± 0.365
2.509GlnVal: 2.509 ± 0.434
1.075GlnTrp: 1.075 ± 0.236
0.777GlnTyr: 0.777 ± 0.203
0.0GlnXaa: 0.0 ± 0.0
Arg
8.185ArgAla: 8.185 ± 0.745
0.956ArgCys: 0.956 ± 0.271
4.122ArgAsp: 4.122 ± 0.486
4.302ArgGlu: 4.302 ± 0.656
2.748ArgPhe: 2.748 ± 0.44
4.6ArgGly: 4.6 ± 0.521
1.733ArgHis: 1.733 ± 0.358
3.525ArgIle: 3.525 ± 0.477
3.226ArgLys: 3.226 ± 0.439
5.616ArgLeu: 5.616 ± 0.536
2.45ArgMet: 2.45 ± 0.33
2.629ArgAsn: 2.629 ± 0.394
4.481ArgPro: 4.481 ± 0.581
2.987ArgGln: 2.987 ± 0.455
6.751ArgArg: 6.751 ± 0.76
4.242ArgSer: 4.242 ± 0.535
4.361ArgThr: 4.361 ± 0.625
5.257ArgVal: 5.257 ± 0.72
2.091ArgTrp: 2.091 ± 0.368
2.151ArgTyr: 2.151 ± 0.403
0.0ArgXaa: 0.0 ± 0.0
Ser
7.827SerAla: 7.827 ± 0.848
0.119SerCys: 0.119 ± 0.088
4.063SerAsp: 4.063 ± 0.37
3.107SerGlu: 3.107 ± 0.426
1.434SerPhe: 1.434 ± 0.305
6.632SerGly: 6.632 ± 0.774
0.657SerHis: 0.657 ± 0.182
2.091SerIle: 2.091 ± 0.317
1.494SerLys: 1.494 ± 0.302
4.122SerLeu: 4.122 ± 0.543
1.673SerMet: 1.673 ± 0.324
1.195SerAsn: 1.195 ± 0.295
2.987SerPro: 2.987 ± 0.377
2.091SerGln: 2.091 ± 0.354
3.465SerArg: 3.465 ± 0.572
3.525SerSer: 3.525 ± 0.477
3.764SerThr: 3.764 ± 0.43
3.525SerVal: 3.525 ± 0.459
1.195SerTrp: 1.195 ± 0.205
1.075SerTyr: 1.075 ± 0.25
0.0SerXaa: 0.0 ± 0.0
Thr
9.738ThrAla: 9.738 ± 0.737
0.896ThrCys: 0.896 ± 0.273
4.302ThrAsp: 4.302 ± 0.402
3.107ThrGlu: 3.107 ± 0.519
1.792ThrPhe: 1.792 ± 0.283
5.735ThrGly: 5.735 ± 0.684
1.195ThrHis: 1.195 ± 0.317
4.003ThrIle: 4.003 ± 0.433
1.852ThrLys: 1.852 ± 0.355
6.333ThrLeu: 6.333 ± 0.701
0.657ThrMet: 0.657 ± 0.19
2.031ThrAsn: 2.031 ± 0.375
4.66ThrPro: 4.66 ± 0.67
1.792ThrGln: 1.792 ± 0.346
4.541ThrArg: 4.541 ± 0.593
3.704ThrSer: 3.704 ± 0.584
5.257ThrThr: 5.257 ± 0.617
5.198ThrVal: 5.198 ± 0.66
1.434ThrTrp: 1.434 ± 0.293
1.075ThrTyr: 1.075 ± 0.273
0.0ThrXaa: 0.0 ± 0.0
Val
7.05ValAla: 7.05 ± 0.544
0.896ValCys: 0.896 ± 0.224
5.496ValAsp: 5.496 ± 0.622
3.883ValGlu: 3.883 ± 0.585
1.553ValPhe: 1.553 ± 0.301
5.377ValGly: 5.377 ± 0.465
1.016ValHis: 1.016 ± 0.254
3.047ValIle: 3.047 ± 0.476
2.27ValLys: 2.27 ± 0.396
4.361ValLeu: 4.361 ± 0.509
1.374ValMet: 1.374 ± 0.362
2.091ValAsn: 2.091 ± 0.381
4.182ValPro: 4.182 ± 0.544
2.39ValGln: 2.39 ± 0.357
4.839ValArg: 4.839 ± 0.543
4.481ValSer: 4.481 ± 0.58
5.556ValThr: 5.556 ± 0.473
4.959ValVal: 4.959 ± 0.581
1.374ValTrp: 1.374 ± 0.327
1.016ValTyr: 1.016 ± 0.218
0.0ValXaa: 0.0 ± 0.0
Trp
2.688TrpAla: 2.688 ± 0.502
0.179TrpCys: 0.179 ± 0.104
0.956TrpAsp: 0.956 ± 0.229
1.075TrpGlu: 1.075 ± 0.227
0.597TrpPhe: 0.597 ± 0.206
1.374TrpGly: 1.374 ± 0.231
0.777TrpHis: 0.777 ± 0.246
0.896TrpIle: 0.896 ± 0.226
0.597TrpLys: 0.597 ± 0.172
2.211TrpLeu: 2.211 ± 0.378
0.836TrpMet: 0.836 ± 0.193
0.836TrpAsn: 0.836 ± 0.226
0.538TrpPro: 0.538 ± 0.218
1.016TrpGln: 1.016 ± 0.262
1.912TrpArg: 1.912 ± 0.375
1.075TrpSer: 1.075 ± 0.231
1.314TrpThr: 1.314 ± 0.239
1.673TrpVal: 1.673 ± 0.333
0.836TrpTrp: 0.836 ± 0.308
0.418TrpTyr: 0.418 ± 0.129
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.27TyrAla: 2.27 ± 0.305
0.239TyrCys: 0.239 ± 0.119
1.852TyrAsp: 1.852 ± 0.257
1.135TyrGlu: 1.135 ± 0.216
0.717TyrPhe: 0.717 ± 0.181
1.613TyrGly: 1.613 ± 0.274
0.299TyrHis: 0.299 ± 0.156
0.836TyrIle: 0.836 ± 0.224
0.538TyrLys: 0.538 ± 0.193
1.553TyrLeu: 1.553 ± 0.296
0.358TyrMet: 0.358 ± 0.12
0.538TyrAsn: 0.538 ± 0.148
0.896TyrPro: 0.896 ± 0.274
0.836TyrGln: 0.836 ± 0.201
1.912TyrArg: 1.912 ± 0.416
1.434TyrSer: 1.434 ± 0.312
1.494TyrThr: 1.494 ± 0.274
1.673TyrVal: 1.673 ± 0.343
0.418TyrTrp: 0.418 ± 0.184
0.358TyrTyr: 0.358 ± 0.144
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 76 proteins (16739 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski