Amino acid dipepetide frequency for Grapevine badnavirus 1

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
3.759AlaAla: 3.759 ± 1.628
0.94AlaCys: 0.94 ± 0.459
1.88AlaAsp: 1.88 ± 1.393
4.229AlaGlu: 4.229 ± 2.063
1.41AlaPhe: 1.41 ± 0.688
2.35AlaGly: 2.35 ± 1.286
0.47AlaHis: 0.47 ± 0.229
3.289AlaIle: 3.289 ± 1.857
2.35AlaLys: 2.35 ± 1.415
5.169AlaLeu: 5.169 ± 3.045
2.82AlaMet: 2.82 ± 1.376
1.88AlaAsn: 1.88 ± 0.917
3.759AlaPro: 3.759 ± 1.256
2.35AlaGln: 2.35 ± 2.315
5.169AlaArg: 5.169 ± 2.522
2.82AlaSer: 2.82 ± 1.376
4.699AlaThr: 4.699 ± 3.035
5.169AlaVal: 5.169 ± 2.522
0.0AlaTrp: 0.0 ± 0.0
2.82AlaTyr: 2.82 ± 1.376
0.0AlaXaa: 0.0 ± 0.0
Cys
0.47CysAla: 0.47 ± 0.229
0.47CysCys: 0.47 ± 0.229
0.47CysAsp: 0.47 ± 0.229
0.47CysGlu: 0.47 ± 0.229
0.47CysPhe: 0.47 ± 0.229
0.94CysGly: 0.94 ± 0.459
0.0CysHis: 0.0 ± 0.0
0.47CysIle: 0.47 ± 0.229
2.82CysLys: 2.82 ± 1.376
1.41CysLeu: 1.41 ± 1.528
0.47CysMet: 0.47 ± 0.229
0.94CysAsn: 0.94 ± 0.459
1.88CysPro: 1.88 ± 1.393
2.82CysGln: 2.82 ± 1.212
0.94CysArg: 0.94 ± 1.683
0.0CysSer: 0.0 ± 0.0
0.0CysThr: 0.0 ± 0.0
0.47CysVal: 0.47 ± 0.229
0.0CysTrp: 0.0 ± 0.0
0.94CysTyr: 0.94 ± 0.459
0.0CysXaa: 0.0 ± 0.0
Asp
2.82AspAla: 2.82 ± 1.376
0.47AspCys: 0.47 ± 0.229
3.289AspAsp: 3.289 ± 1.271
5.169AspGlu: 5.169 ± 1.459
2.35AspPhe: 2.35 ± 1.146
3.759AspGly: 3.759 ± 1.191
1.88AspHis: 1.88 ± 1.393
4.699AspIle: 4.699 ± 1.17
2.35AspLys: 2.35 ± 1.146
4.229AspLeu: 4.229 ± 1.283
0.47AspMet: 0.47 ± 0.229
2.35AspAsn: 2.35 ± 1.146
2.35AspPro: 2.35 ± 1.415
2.35AspGln: 2.35 ± 1.146
1.88AspArg: 1.88 ± 1.393
0.94AspSer: 0.94 ± 0.459
2.35AspThr: 2.35 ± 1.146
1.88AspVal: 1.88 ± 1.393
0.47AspTrp: 0.47 ± 0.229
3.759AspTyr: 3.759 ± 1.256
0.0AspXaa: 0.0 ± 0.0
Glu
6.109GluAla: 6.109 ± 1.724
1.41GluCys: 1.41 ± 1.528
7.989GluAsp: 7.989 ± 3.897
12.218GluGlu: 12.218 ± 2.812
1.41GluPhe: 1.41 ± 1.528
6.579GluGly: 6.579 ± 0.262
1.88GluHis: 1.88 ± 1.393
4.699GluIle: 4.699 ± 2.571
7.519GluLys: 7.519 ± 4.145
5.639GluLeu: 5.639 ± 2.613
0.47GluMet: 0.47 ± 0.229
4.699GluAsn: 4.699 ± 1.337
0.94GluPro: 0.94 ± 1.683
5.169GluGln: 5.169 ± 2.522
4.229GluArg: 4.229 ± 1.245
7.989GluSer: 7.989 ± 8.469
1.88GluThr: 1.88 ± 1.393
5.169GluVal: 5.169 ± 2.522
0.94GluTrp: 0.94 ± 0.459
0.94GluTyr: 0.94 ± 0.459
0.0GluXaa: 0.0 ± 0.0
Phe
2.35PheAla: 2.35 ± 1.146
0.47PheCys: 0.47 ± 0.229
0.47PheAsp: 0.47 ± 0.229
2.35PheGlu: 2.35 ± 1.146
0.94PhePhe: 0.94 ± 0.459
0.47PheGly: 0.47 ± 0.229
1.41PheHis: 1.41 ± 0.688
3.759PheIle: 3.759 ± 1.191
2.82PheLys: 2.82 ± 1.325
2.35PheLeu: 2.35 ± 1.146
0.47PheMet: 0.47 ± 0.229
2.35PheAsn: 2.35 ± 1.146
0.47PhePro: 0.47 ± 0.229
1.41PheGln: 1.41 ± 0.688
2.35PheArg: 2.35 ± 1.286
2.82PheSer: 2.82 ± 1.212
0.94PheThr: 0.94 ± 0.459
2.35PheVal: 2.35 ± 1.415
0.47PheTrp: 0.47 ± 0.229
1.41PheTyr: 1.41 ± 0.688
0.0PheXaa: 0.0 ± 0.0
Gly
2.35GlyAla: 2.35 ± 1.286
1.41GlyCys: 1.41 ± 0.688
1.41GlyAsp: 1.41 ± 0.688
7.049GlyGlu: 7.049 ± 2.01
2.35GlyPhe: 2.35 ± 1.415
4.229GlyGly: 4.229 ± 1.399
1.41GlyHis: 1.41 ± 3.534
4.699GlyIle: 4.699 ± 3.253
3.759GlyLys: 3.759 ± 1.834
2.82GlyLeu: 2.82 ± 1.376
2.82GlyMet: 2.82 ± 1.309
3.289GlyAsn: 3.289 ± 2.916
0.94GlyPro: 0.94 ± 0.459
3.289GlyGln: 3.289 ± 1.271
4.229GlyArg: 4.229 ± 2.671
1.88GlySer: 1.88 ± 1.535
3.759GlyThr: 3.759 ± 1.256
5.169GlyVal: 5.169 ± 2.522
1.88GlyTrp: 1.88 ± 0.917
1.88GlyTyr: 1.88 ± 0.917
0.0GlyXaa: 0.0 ± 0.0
His
1.41HisAla: 1.41 ± 1.528
1.88HisCys: 1.88 ± 1.393
1.88HisAsp: 1.88 ± 1.393
1.41HisGlu: 1.41 ± 1.528
0.47HisPhe: 0.47 ± 0.229
0.47HisGly: 0.47 ± 1.854
1.41HisHis: 1.41 ± 1.528
1.41HisIle: 1.41 ± 0.688
0.47HisLys: 0.47 ± 0.229
2.82HisLeu: 2.82 ± 1.212
0.0HisMet: 0.0 ± 0.0
0.47HisAsn: 0.47 ± 0.229
0.94HisPro: 0.94 ± 0.459
0.94HisGln: 0.94 ± 0.459
1.88HisArg: 1.88 ± 0.917
0.0HisSer: 0.0 ± 0.0
0.94HisThr: 0.94 ± 1.683
1.88HisVal: 1.88 ± 0.917
0.47HisTrp: 0.47 ± 0.229
0.0HisTyr: 0.0 ± 0.0
0.0HisXaa: 0.0 ± 0.0
Ile
2.82IleAla: 2.82 ± 1.376
1.41IleCys: 1.41 ± 0.688
2.82IleAsp: 2.82 ± 1.376
5.169IleGlu: 5.169 ± 0.942
0.47IlePhe: 0.47 ± 0.229
2.82IleGly: 2.82 ± 1.376
1.41IleHis: 1.41 ± 1.528
3.289IleIle: 3.289 ± 1.605
4.699IleLys: 4.699 ± 6.58
4.229IleLeu: 4.229 ± 3.463
1.88IleMet: 1.88 ± 0.917
4.229IleAsn: 4.229 ± 2.063
5.639IlePro: 5.639 ± 1.605
6.109IleGln: 6.109 ± 1.724
4.699IleArg: 4.699 ± 1.349
5.169IleSer: 5.169 ± 3.045
3.289IleThr: 3.289 ± 2.916
2.35IleVal: 2.35 ± 1.146
0.47IleTrp: 0.47 ± 0.229
2.82IleTyr: 2.82 ± 1.376
0.0IleXaa: 0.0 ± 0.0
Lys
2.35LysAla: 2.35 ± 3.51
1.41LysCys: 1.41 ± 0.688
4.229LysAsp: 4.229 ± 2.944
7.989LysGlu: 7.989 ± 3.027
2.35LysPhe: 2.35 ± 1.286
3.759LysGly: 3.759 ± 1.256
1.88LysHis: 1.88 ± 0.917
3.289LysIle: 3.289 ± 1.605
4.229LysLys: 4.229 ± 2.063
7.989LysLeu: 7.989 ± 3.027
4.229LysMet: 4.229 ± 1.371
3.759LysAsn: 3.759 ± 1.834
2.82LysPro: 2.82 ± 3.354
0.47LysGln: 0.47 ± 2.01
5.639LysArg: 5.639 ± 1.576
5.639LysSer: 5.639 ± 1.605
1.41LysThr: 1.41 ± 1.528
2.82LysVal: 2.82 ± 1.325
0.94LysTrp: 0.94 ± 0.459
0.94LysTyr: 0.94 ± 0.459
0.0LysXaa: 0.0 ± 0.0
Leu
2.35LeuAla: 2.35 ± 1.286
1.88LeuCys: 1.88 ± 0.917
2.35LeuAsp: 2.35 ± 1.286
7.989LeuGlu: 7.989 ± 1.646
1.41LeuPhe: 1.41 ± 1.528
7.049LeuGly: 7.049 ± 2.517
0.47LeuHis: 0.47 ± 0.229
2.35LeuIle: 2.35 ± 1.286
9.398LeuLys: 9.398 ± 2.341
4.229LeuLeu: 4.229 ± 2.063
0.94LeuMet: 0.94 ± 1.16
2.82LeuAsn: 2.82 ± 1.325
3.289LeuPro: 3.289 ± 1.271
4.699LeuGln: 4.699 ± 1.349
5.169LeuArg: 5.169 ± 8.43
7.049LeuSer: 7.049 ± 2.01
3.289LeuThr: 3.289 ± 1.605
7.519LeuVal: 7.519 ± 2.076
1.88LeuTrp: 1.88 ± 0.917
3.759LeuTyr: 3.759 ± 1.256
0.0LeuXaa: 0.0 ± 0.0
Met
0.94MetAla: 0.94 ± 0.459
0.0MetCys: 0.0 ± 0.0
1.41MetAsp: 1.41 ± 0.688
3.759MetGlu: 3.759 ± 1.191
2.82MetPhe: 2.82 ± 1.376
1.41MetGly: 1.41 ± 0.688
0.47MetHis: 0.47 ± 0.229
1.88MetIle: 1.88 ± 0.917
2.82MetLys: 2.82 ± 1.376
2.35MetLeu: 2.35 ± 1.146
0.47MetMet: 0.47 ± 0.229
2.35MetAsn: 2.35 ± 1.146
0.94MetPro: 0.94 ± 0.459
1.41MetGln: 1.41 ± 0.688
0.94MetArg: 0.94 ± 0.459
0.94MetSer: 0.94 ± 1.837
1.88MetThr: 1.88 ± 0.917
1.41MetVal: 1.41 ± 0.688
0.0MetTrp: 0.0 ± 0.0
0.47MetTyr: 0.47 ± 0.229
0.0MetXaa: 0.0 ± 0.0
Asn
1.88AsnAla: 1.88 ± 0.917
0.0AsnCys: 0.0 ± 0.0
1.41AsnAsp: 1.41 ± 0.688
1.41AsnGlu: 1.41 ± 0.688
2.82AsnPhe: 2.82 ± 1.376
2.35AsnGly: 2.35 ± 1.286
1.41AsnHis: 1.41 ± 0.688
2.82AsnIle: 2.82 ± 1.376
3.759AsnLys: 3.759 ± 1.834
6.579AsnLeu: 6.579 ± 0.262
0.94AsnMet: 0.94 ± 0.459
1.41AsnAsn: 1.41 ± 4.762
2.82AsnPro: 2.82 ± 1.376
2.82AsnGln: 2.82 ± 1.376
2.35AsnArg: 2.35 ± 1.146
4.229AsnSer: 4.229 ± 1.245
3.289AsnThr: 3.289 ± 1.857
0.94AsnVal: 0.94 ± 0.459
1.41AsnTrp: 1.41 ± 1.528
1.88AsnTyr: 1.88 ± 2.544
0.0AsnXaa: 0.0 ± 0.0
Pro
3.289ProAla: 3.289 ± 1.271
0.0ProCys: 0.0 ± 0.0
3.289ProAsp: 3.289 ± 1.605
3.289ProGlu: 3.289 ± 1.271
0.94ProPhe: 0.94 ± 0.459
4.699ProGly: 4.699 ± 4.442
0.94ProHis: 0.94 ± 0.459
2.35ProIle: 2.35 ± 1.146
2.35ProLys: 2.35 ± 1.146
2.35ProLeu: 2.35 ± 1.415
0.94ProMet: 0.94 ± 0.459
2.82ProAsn: 2.82 ± 1.212
4.229ProPro: 4.229 ± 2.063
0.94ProGln: 0.94 ± 0.459
0.94ProArg: 0.94 ± 0.459
4.229ProSer: 4.229 ± 3.463
3.759ProThr: 3.759 ± 1.256
2.82ProVal: 2.82 ± 1.376
0.47ProTrp: 0.47 ± 0.229
0.94ProTyr: 0.94 ± 0.459
0.0ProXaa: 0.0 ± 0.0
Gln
2.82GlnAla: 2.82 ± 1.376
0.94GlnCys: 0.94 ± 0.459
1.41GlnAsp: 1.41 ± 0.688
5.639GlnGlu: 5.639 ± 2.751
1.41GlnPhe: 1.41 ± 0.688
2.82GlnGly: 2.82 ± 2.086
1.88GlnHis: 1.88 ± 1.393
6.579GlnIle: 6.579 ± 4.357
2.82GlnLys: 2.82 ± 1.212
5.639GlnLeu: 5.639 ± 4.172
0.94GlnMet: 0.94 ± 0.459
2.82GlnAsn: 2.82 ± 1.325
2.35GlnPro: 2.35 ± 1.286
4.229GlnGln: 4.229 ± 1.245
2.35GlnArg: 2.35 ± 1.286
3.759GlnSer: 3.759 ± 1.256
1.88GlnThr: 1.88 ± 0.917
1.41GlnVal: 1.41 ± 0.688
0.94GlnTrp: 0.94 ± 0.459
1.88GlnTyr: 1.88 ± 0.917
0.0GlnXaa: 0.0 ± 0.0
Arg
5.169ArgAla: 5.169 ± 2.488
0.47ArgCys: 0.47 ± 0.229
5.169ArgAsp: 5.169 ± 2.823
2.82ArgGlu: 2.82 ± 3.354
2.35ArgPhe: 2.35 ± 1.146
3.289ArgGly: 3.289 ± 1.857
1.88ArgHis: 1.88 ± 0.917
4.699ArgIle: 4.699 ± 1.349
4.229ArgLys: 4.229 ± 1.245
4.229ArgLeu: 4.229 ± 2.063
3.289ArgMet: 3.289 ± 1.605
3.289ArgAsn: 3.289 ± 1.179
1.88ArgPro: 1.88 ± 0.917
1.88ArgGln: 1.88 ± 1.535
5.639ArgArg: 5.639 ± 2.751
3.759ArgSer: 3.759 ± 2.787
2.35ArgThr: 2.35 ± 1.146
3.759ArgVal: 3.759 ± 1.628
1.88ArgTrp: 1.88 ± 0.917
2.35ArgTyr: 2.35 ± 1.146
0.0ArgXaa: 0.0 ± 0.0
Ser
2.82SerAla: 2.82 ± 3.354
1.41SerCys: 1.41 ± 1.528
2.35SerAsp: 2.35 ± 2.315
6.109SerGlu: 6.109 ± 0.486
3.759SerPhe: 3.759 ± 1.834
4.229SerGly: 4.229 ± 2.063
1.88SerHis: 1.88 ± 1.393
3.759SerIle: 3.759 ± 1.256
3.289SerLys: 3.289 ± 3.89
8.459SerLeu: 8.459 ± 6.259
0.94SerMet: 0.94 ± 0.459
1.88SerAsn: 1.88 ± 2.544
3.289SerPro: 3.289 ± 1.857
3.759SerGln: 3.759 ± 1.628
5.639SerArg: 5.639 ± 1.576
5.169SerSer: 5.169 ± 6.357
4.229SerThr: 4.229 ± 2.063
1.41SerVal: 1.41 ± 0.688
1.41SerTrp: 1.41 ± 0.688
2.82SerTyr: 2.82 ± 1.212
0.0SerXaa: 0.0 ± 0.0
Thr
5.639ThrAla: 5.639 ± 0.713
0.47ThrCys: 0.47 ± 0.229
3.289ThrAsp: 3.289 ± 1.605
3.759ThrGlu: 3.759 ± 1.628
0.47ThrPhe: 0.47 ± 0.229
5.169ThrGly: 5.169 ± 2.522
0.0ThrHis: 0.0 ± 0.0
2.82ThrIle: 2.82 ± 3.056
0.94ThrLys: 0.94 ± 0.459
2.82ThrLeu: 2.82 ± 1.376
2.35ThrMet: 2.35 ± 1.146
0.47ThrAsn: 0.47 ± 0.229
3.289ThrPro: 3.289 ± 1.605
4.229ThrGln: 4.229 ± 3.25
2.35ThrArg: 2.35 ± 1.286
2.82ThrSer: 2.82 ± 1.376
2.35ThrThr: 2.35 ± 1.286
3.289ThrVal: 3.289 ± 1.857
0.0ThrTrp: 0.0 ± 0.0
1.88ThrTyr: 1.88 ± 1.535
0.0ThrXaa: 0.0 ± 0.0
Val
4.699ValAla: 4.699 ± 1.337
0.94ValCys: 0.94 ± 1.683
1.88ValAsp: 1.88 ± 0.917
1.88ValGlu: 1.88 ± 1.535
3.759ValPhe: 3.759 ± 1.256
3.289ValGly: 3.289 ± 1.271
0.47ValHis: 0.47 ± 0.229
4.699ValIle: 4.699 ± 1.349
1.41ValLys: 1.41 ± 1.677
1.88ValLeu: 1.88 ± 0.917
2.82ValMet: 2.82 ± 1.376
2.82ValAsn: 2.82 ± 1.376
2.35ValPro: 2.35 ± 1.146
3.759ValGln: 3.759 ± 2.787
4.699ValArg: 4.699 ± 2.293
3.289ValSer: 3.289 ± 1.605
4.229ValThr: 4.229 ± 1.283
1.88ValVal: 1.88 ± 0.917
0.0ValTrp: 0.0 ± 0.0
2.82ValTyr: 2.82 ± 1.376
0.0ValXaa: 0.0 ± 0.0
Trp
0.0TrpAla: 0.0 ± 0.0
0.47TrpCys: 0.47 ± 0.229
0.47TrpAsp: 0.47 ± 0.229
2.35TrpGlu: 2.35 ± 1.286
0.0TrpPhe: 0.0 ± 0.0
0.47TrpGly: 0.47 ± 0.229
0.0TrpHis: 0.0 ± 0.0
0.94TrpIle: 0.94 ± 0.459
2.35TrpLys: 2.35 ± 1.146
1.88TrpLeu: 1.88 ± 0.917
0.47TrpMet: 0.47 ± 0.229
0.0TrpAsn: 0.0 ± 0.0
0.0TrpPro: 0.0 ± 0.0
1.41TrpGln: 1.41 ± 0.688
1.41TrpArg: 1.41 ± 0.688
1.41TrpSer: 1.41 ± 1.528
0.47TrpThr: 0.47 ± 0.229
0.47TrpVal: 0.47 ± 0.229
0.0TrpTrp: 0.0 ± 0.0
0.0TrpTyr: 0.0 ± 0.0
0.0TrpXaa: 0.0 ± 0.0
Tyr
3.289TyrAla: 3.289 ± 1.605
0.0TyrCys: 0.0 ± 0.0
2.35TyrAsp: 2.35 ± 1.146
2.82TyrGlu: 2.82 ± 1.376
0.47TyrPhe: 0.47 ± 0.229
0.47TyrGly: 0.47 ± 0.229
0.0TyrHis: 0.0 ± 0.0
3.289TyrIle: 3.289 ± 1.605
3.759TyrLys: 3.759 ± 3.069
3.289TyrLeu: 3.289 ± 1.271
0.47TyrMet: 0.47 ± 0.229
1.88TyrAsn: 1.88 ± 0.917
1.41TyrPro: 1.41 ± 0.688
0.47TyrGln: 0.47 ± 0.229
1.88TyrArg: 1.88 ± 0.917
4.699TyrSer: 4.699 ± 1.17
1.41TyrThr: 1.41 ± 0.688
1.41TyrVal: 1.41 ± 0.688
0.94TyrTrp: 0.94 ± 1.683
1.41TyrTyr: 1.41 ± 0.688
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 3 proteins (2129 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski