Amino acid dipepetide frequency for Cebus imitator (Panamanian white-faced capuchin) (Cebus capucinus imitator)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	6.893AlaAla: 6.893 ± 0.04	1.37AlaCys: 1.37 ± 0.01	2.955AlaAsp: 2.955 ± 0.013	4.838AlaGlu: 4.838 ± 0.025	2.612AlaPhe: 2.612 ± 0.014	4.73AlaGly: 4.73 ± 0.022	1.54AlaHis: 1.54 ± 0.008	2.78AlaIle: 2.78 ± 0.013	3.511AlaLys: 3.511 ± 0.02	7.084AlaLeu: 7.084 ± 0.029	1.522AlaMet: 1.522 ± 0.011	2.031AlaAsn: 2.031 ± 0.01	4.112AlaPro: 4.112 ± 0.027	3.267AlaGln: 3.267 ± 0.018	3.636AlaArg: 3.636 ± 0.018	5.705AlaSer: 5.705 ± 0.022	3.632AlaThr: 3.632 ± 0.015	4.684AlaVal: 4.684 ± 0.018	0.795AlaTrp: 0.795 ± 0.008	1.546AlaTyr: 1.546 ± 0.01	0.0AlaXaa: 0.0 ± 0.0
Cys	1.202CysAla: 1.202 ± 0.01	0.64CysCys: 0.64 ± 0.009	0.985CysAsp: 0.985 ± 0.011	1.283CysGlu: 1.283 ± 0.012	0.833CysPhe: 0.833 ± 0.007	1.712CysGly: 1.712 ± 0.016	0.66CysHis: 0.66 ± 0.007	0.95CysIle: 0.95 ± 0.009	1.171CysLys: 1.171 ± 0.009	2.149CysLeu: 2.149 ± 0.013	0.415CysMet: 0.415 ± 0.005	0.818CysAsn: 0.818 ± 0.009	1.333CysPro: 1.333 ± 0.013	1.058CysGln: 1.058 ± 0.01	1.275CysArg: 1.275 ± 0.01	1.958CysSer: 1.958 ± 0.014	1.081CysThr: 1.081 ± 0.009	1.283CysVal: 1.283 ± 0.01	0.286CysTrp: 0.286 ± 0.004	0.566CysTyr: 0.566 ± 0.005	0.0CysXaa: 0.0 ± 0.0
Asp	2.871AspAla: 2.871 ± 0.013	1.052AspCys: 1.052 ± 0.009	2.688AspAsp: 2.688 ± 0.015	3.526AspGlu: 3.526 ± 0.017	2.153AspPhe: 2.153 ± 0.012	3.335AspGly: 3.335 ± 0.019	1.152AspHis: 1.152 ± 0.007	2.659AspIle: 2.659 ± 0.014	2.578AspLys: 2.578 ± 0.012	5.042AspLeu: 5.042 ± 0.019	1.127AspMet: 1.127 ± 0.008	1.696AspAsn: 1.696 ± 0.01	2.919AspPro: 2.919 ± 0.014	1.854AspGln: 1.854 ± 0.01	2.423AspArg: 2.423 ± 0.012	4.2AspSer: 4.2 ± 0.02	2.482AspThr: 2.482 ± 0.014	3.091AspVal: 3.091 ± 0.016	0.623AspTrp: 0.623 ± 0.006	1.515AspTyr: 1.515 ± 0.01	0.0AspXaa: 0.0 ± 0.0
Glu	5.307GluAla: 5.307 ± 0.026	1.459GluCys: 1.459 ± 0.018	4.529GluAsp: 4.529 ± 0.019	8.086GluGlu: 8.086 ± 0.044	2.08GluPhe: 2.08 ± 0.011	4.2GluGly: 4.2 ± 0.018	1.541GluHis: 1.541 ± 0.009	3.2GluIle: 3.2 ± 0.018	5.545GluLys: 5.545 ± 0.03	6.574GluLeu: 6.574 ± 0.029	1.713GluMet: 1.713 ± 0.011	3.2GluAsn: 3.2 ± 0.017	3.241GluPro: 3.241 ± 0.022	3.188GluGln: 3.188 ± 0.017	4.044GluArg: 4.044 ± 0.021	4.402GluSer: 4.402 ± 0.018	3.424GluThr: 3.424 ± 0.018	4.203GluVal: 4.203 ± 0.018	0.709GluTrp: 0.709 ± 0.006	1.637GluTyr: 1.637 ± 0.013	0.0GluXaa: 0.0 ± 0.0
Phe	1.986PheAla: 1.986 ± 0.011	0.905PheCys: 0.905 ± 0.007	1.71PheAsp: 1.71 ± 0.009	2.041PheGlu: 2.041 ± 0.011	1.561PhePhe: 1.561 ± 0.013	2.204PheGly: 2.204 ± 0.014	1.05PheHis: 1.05 ± 0.007	1.819PheIle: 1.819 ± 0.013	1.823PheLys: 1.823 ± 0.011	4.006PheLeu: 4.006 ± 0.02	0.772PheMet: 0.772 ± 0.006	1.357PheAsn: 1.357 ± 0.01	1.982PhePro: 1.982 ± 0.011	1.813PheGln: 1.813 ± 0.01	1.99PheArg: 1.99 ± 0.013	3.374PheSer: 3.374 ± 0.015	2.029PheThr: 2.029 ± 0.013	2.111PheVal: 2.111 ± 0.012	0.488PheTrp: 0.488 ± 0.005	1.163PheTyr: 1.163 ± 0.01	0.001PheXaa: 0.001 ± 0.0
Gly	4.491GlyAla: 4.491 ± 0.025	1.25GlyCys: 1.25 ± 0.01	3.166GlyAsp: 3.166 ± 0.014	4.187GlyGlu: 4.187 ± 0.021	2.398GlyPhe: 2.398 ± 0.016	5.104GlyGly: 5.104 ± 0.035	1.685GlyHis: 1.685 ± 0.011	2.79GlyIle: 2.79 ± 0.014	3.889GlyLys: 3.889 ± 0.018	5.857GlyLeu: 5.857 ± 0.025	1.321GlyMet: 1.321 ± 0.01	2.359GlyAsn: 2.359 ± 0.013	4.23GlyPro: 4.23 ± 0.04	2.736GlyGln: 2.736 ± 0.014	3.732GlyArg: 3.732 ± 0.018	5.799GlySer: 5.799 ± 0.026	3.578GlyThr: 3.578 ± 0.015	3.52GlyVal: 3.52 ± 0.017	0.793GlyTrp: 0.793 ± 0.007	1.758GlyTyr: 1.758 ± 0.011	0.0GlyXaa: 0.0 ± 0.0
His	1.329HisAla: 1.329 ± 0.008	0.726HisCys: 0.726 ± 0.008	0.91HisAsp: 0.91 ± 0.006	1.339HisGlu: 1.339 ± 0.009	1.087HisPhe: 1.087 ± 0.007	1.534HisGly: 1.534 ± 0.01	0.917HisHis: 0.917 ± 0.008	1.27HisIle: 1.27 ± 0.01	1.297HisLys: 1.297 ± 0.01	2.923HisLeu: 2.923 ± 0.013	0.596HisMet: 0.596 ± 0.006	0.88HisAsn: 0.88 ± 0.007	1.65HisPro: 1.65 ± 0.011	1.361HisGln: 1.361 ± 0.012	1.588HisArg: 1.588 ± 0.01	2.327HisSer: 2.327 ± 0.013	1.526HisThr: 1.526 ± 0.013	1.49HisVal: 1.49 ± 0.008	0.351HisTrp: 0.351 ± 0.004	0.804HisTyr: 0.804 ± 0.006	0.0HisXaa: 0.0 ± 0.0
Ile	2.6IleAla: 2.6 ± 0.012	1.062IleCys: 1.062 ± 0.009	2.059IleAsp: 2.059 ± 0.012	2.671IleGlu: 2.671 ± 0.016	1.879IlePhe: 1.879 ± 0.012	2.24IleGly: 2.24 ± 0.014	1.386IleHis: 1.386 ± 0.011	2.388IleIle: 2.388 ± 0.017	2.703IleLys: 2.703 ± 0.018	4.534IleLeu: 4.534 ± 0.019	1.007IleMet: 1.007 ± 0.007	1.861IleAsn: 1.861 ± 0.013	2.603IlePro: 2.603 ± 0.012	2.306IleGln: 2.306 ± 0.012	2.399IleArg: 2.399 ± 0.01	3.704IleSer: 3.704 ± 0.016	2.575IleThr: 2.575 ± 0.019	2.504IleVal: 2.504 ± 0.015	0.503IleTrp: 0.503 ± 0.005	1.377IleTyr: 1.377 ± 0.011	0.0IleXaa: 0.0 ± 0.0
Lys	4.096LysAla: 4.096 ± 0.022	1.167LysCys: 1.167 ± 0.011	3.215LysAsp: 3.215 ± 0.018	5.222LysGlu: 5.222 ± 0.028	1.769LysPhe: 1.769 ± 0.01	3.312LysGly: 3.312 ± 0.02	1.432LysHis: 1.432 ± 0.009	2.844LysIle: 2.844 ± 0.017	4.824LysLys: 4.824 ± 0.026	5.279LysLeu: 5.279 ± 0.023	1.463LysMet: 1.463 ± 0.01	2.465LysAsn: 2.465 ± 0.012	3.169LysPro: 3.169 ± 0.021	2.674LysGln: 2.674 ± 0.014	3.361LysArg: 3.361 ± 0.017	3.924LysSer: 3.924 ± 0.019	3.138LysThr: 3.138 ± 0.017	3.505LysVal: 3.505 ± 0.018	0.607LysTrp: 0.607 ± 0.007	1.608LysTyr: 1.608 ± 0.012	0.0LysXaa: 0.0 ± 0.0
Leu	6.776LeuAla: 6.776 ± 0.029	2.163LeuCys: 2.163 ± 0.013	4.75LeuAsp: 4.75 ± 0.019	7.286LeuGlu: 7.286 ± 0.031	3.34LeuPhe: 3.34 ± 0.018	5.839LeuGly: 5.839 ± 0.024	2.73LeuHis: 2.73 ± 0.015	3.866LeuIle: 3.866 ± 0.019	5.901LeuLys: 5.901 ± 0.026	10.703LeuLeu: 10.703 ± 0.047	2.031LeuMet: 2.031 ± 0.012	3.523LeuAsn: 3.523 ± 0.016	5.941LeuPro: 5.941 ± 0.024	5.832LeuGln: 5.832 ± 0.029	5.912LeuArg: 5.912 ± 0.026	7.889LeuSer: 7.889 ± 0.021	5.042LeuThr: 5.042 ± 0.017	5.461LeuVal: 5.461 ± 0.02	1.126LeuTrp: 1.126 ± 0.01	2.534LeuTyr: 2.534 ± 0.013	0.001LeuXaa: 0.001 ± 0.0
Met	1.955MetAla: 1.955 ± 0.009	0.401MetCys: 0.401 ± 0.005	1.273MetAsp: 1.273 ± 0.008	1.918MetGlu: 1.918 ± 0.011	0.729MetPhe: 0.729 ± 0.008	1.305MetGly: 1.305 ± 0.009	0.472MetHis: 0.472 ± 0.005	0.851MetIle: 0.851 ± 0.007	1.473MetLys: 1.473 ± 0.009	1.979MetLeu: 1.979 ± 0.011	0.578MetMet: 0.578 ± 0.006	0.92MetAsn: 0.92 ± 0.008	1.109MetPro: 1.109 ± 0.01	0.941MetGln: 0.941 ± 0.008	1.047MetArg: 1.047 ± 0.008	1.577MetSer: 1.577 ± 0.01	1.132MetThr: 1.132 ± 0.008	1.394MetVal: 1.394 ± 0.008	0.242MetTrp: 0.242 ± 0.003	0.554MetTyr: 0.554 ± 0.005	0.0MetXaa: 0.0 ± 0.0
Asn	2.086AsnAla: 2.086 ± 0.012	0.828AsnCys: 0.828 ± 0.007	1.539AsnAsp: 1.539 ± 0.009	2.271AsnGlu: 2.271 ± 0.013	1.491AsnPhe: 1.491 ± 0.01	2.452AsnGly: 2.452 ± 0.014	0.96AsnHis: 0.96 ± 0.008	2.131AsnIle: 2.131 ± 0.013	2.246AsnLys: 2.246 ± 0.013	3.765AsnLeu: 3.765 ± 0.017	0.938AsnMet: 0.938 ± 0.007	1.533AsnAsn: 1.533 ± 0.012	2.168AsnPro: 2.168 ± 0.013	1.685AsnGln: 1.685 ± 0.011	1.859AsnArg: 1.859 ± 0.01	3.124AsnSer: 3.124 ± 0.015	1.968AsnThr: 1.968 ± 0.011	2.206AsnVal: 2.206 ± 0.013	0.462AsnTrp: 0.462 ± 0.005	1.125AsnTyr: 1.125 ± 0.009	0.0AsnXaa: 0.0 ± 0.0
Pro	4.816ProAla: 4.816 ± 0.028	1.118ProCys: 1.118 ± 0.011	2.783ProAsp: 2.783 ± 0.014	4.459ProGlu: 4.459 ± 0.021	1.887ProPhe: 1.887 ± 0.01	5.276ProGly: 5.276 ± 0.044	1.444ProHis: 1.444 ± 0.01	1.92ProIle: 1.92 ± 0.015	2.805ProLys: 2.805 ± 0.023	5.177ProLeu: 5.177 ± 0.019	1.043ProMet: 1.043 ± 0.009	1.817ProAsn: 1.817 ± 0.011	6.08ProPro: 6.08 ± 0.051	2.827ProGln: 2.827 ± 0.019	3.288ProArg: 3.288 ± 0.02	5.657ProSer: 5.657 ± 0.026	3.101ProThr: 3.101 ± 0.016	3.766ProVal: 3.766 ± 0.019	0.681ProTrp: 0.681 ± 0.007	1.546ProTyr: 1.546 ± 0.013	0.001ProXaa: 0.001 ± 0.0
Gln	3.556GlnAla: 3.556 ± 0.018	0.934GlnCys: 0.934 ± 0.01	2.347GlnAsp: 2.347 ± 0.012	3.951GlnGlu: 3.951 ± 0.023	1.39GlnPhe: 1.39 ± 0.01	2.843GlnGly: 2.843 ± 0.017	1.306GlnHis: 1.306 ± 0.01	2.02GlnIle: 2.02 ± 0.012	2.996GlnLys: 2.996 ± 0.017	4.79GlnLeu: 4.79 ± 0.024	1.129GlnMet: 1.129 ± 0.008	1.865GlnAsn: 1.865 ± 0.011	2.823GlnPro: 2.823 ± 0.017	3.148GlnGln: 3.148 ± 0.029	3.002GlnArg: 3.002 ± 0.019	3.138GlnSer: 3.138 ± 0.014	2.297GlnThr: 2.297 ± 0.011	2.818GlnVal: 2.818 ± 0.014	0.554GlnTrp: 0.554 ± 0.006	1.157GlnTyr: 1.157 ± 0.008	0.0GlnXaa: 0.0 ± 0.0
Arg	3.801ArgAla: 3.801 ± 0.019	1.194ArgCys: 1.194 ± 0.013	2.787ArgAsp: 2.787 ± 0.013	4.0ArgGlu: 4.0 ± 0.02	1.862ArgPhe: 1.862 ± 0.01	3.606ArgGly: 3.606 ± 0.021	1.546ArgHis: 1.546 ± 0.01	2.473ArgIle: 2.473 ± 0.013	3.716ArgLys: 3.716 ± 0.017	5.448ArgLeu: 5.448 ± 0.023	1.209ArgMet: 1.209 ± 0.008	2.118ArgAsn: 2.118 ± 0.01	3.212ArgPro: 3.212 ± 0.017	2.648ArgGln: 2.648 ± 0.015	4.31ArgArg: 4.31 ± 0.024	4.24ArgSer: 4.24 ± 0.024	2.804ArgThr: 2.804 ± 0.014	3.147ArgVal: 3.147 ± 0.018	0.697ArgTrp: 0.697 ± 0.007	1.444ArgTyr: 1.444 ± 0.009	0.0ArgXaa: 0.0 ± 0.0
Ser	5.279SerAla: 5.279 ± 0.02	1.799SerCys: 1.799 ± 0.013	3.896SerAsp: 3.896 ± 0.021	5.213SerGlu: 5.213 ± 0.023	3.012SerPhe: 3.012 ± 0.014	5.617SerGly: 5.617 ± 0.022	2.127SerHis: 2.127 ± 0.013	3.199SerIle: 3.199 ± 0.014	4.136SerLys: 4.136 ± 0.019	8.133SerLeu: 8.133 ± 0.024	1.623SerMet: 1.623 ± 0.009	2.708SerAsn: 2.708 ± 0.015	5.963SerPro: 5.963 ± 0.032	3.889SerGln: 3.889 ± 0.016	4.54SerArg: 4.54 ± 0.021	9.307SerSer: 9.307 ± 0.039	4.405SerThr: 4.405 ± 0.021	4.803SerVal: 4.803 ± 0.016	1.063SerTrp: 1.063 ± 0.009	2.05SerTyr: 2.05 ± 0.012	0.0SerXaa: 0.0 ± 0.0
Thr	3.751ThrAla: 3.751 ± 0.018	1.26ThrCys: 1.26 ± 0.011	2.483ThrAsp: 2.483 ± 0.012	3.542ThrGlu: 3.542 ± 0.019	2.09ThrPhe: 2.09 ± 0.011	3.565ThrGly: 3.565 ± 0.017	1.325ThrHis: 1.325 ± 0.01	2.38ThrIle: 2.38 ± 0.014	2.687ThrLys: 2.687 ± 0.017	5.244ThrLeu: 5.244 ± 0.018	1.116ThrMet: 1.116 ± 0.007	1.752ThrAsn: 1.752 ± 0.009	3.562ThrPro: 3.562 ± 0.019	2.301ThrGln: 2.301 ± 0.012	2.464ThrArg: 2.464 ± 0.011	4.605ThrSer: 4.605 ± 0.018	3.002ThrThr: 3.002 ± 0.024	3.798ThrVal: 3.798 ± 0.019	0.687ThrTrp: 0.687 ± 0.007	1.42ThrTyr: 1.42 ± 0.008	0.0ThrXaa: 0.0 ± 0.0
Val	4.278ValAla: 4.278 ± 0.017	1.404ValCys: 1.404 ± 0.01	2.955ValAsp: 2.955 ± 0.014	3.897ValGlu: 3.897 ± 0.018	2.36ValPhe: 2.36 ± 0.014	3.385ValGly: 3.385 ± 0.013	1.552ValHis: 1.552 ± 0.01	2.899ValIle: 2.899 ± 0.015	3.438ValLys: 3.438 ± 0.018	6.109ValLeu: 6.109 ± 0.023	1.318ValMet: 1.318 ± 0.009	2.278ValAsn: 2.278 ± 0.013	3.572ValPro: 3.572 ± 0.018	2.718ValGln: 2.718 ± 0.012	3.043ValArg: 3.043 ± 0.014	4.795ValSer: 4.795 ± 0.021	3.721ValThr: 3.721 ± 0.024	3.932ValVal: 3.932 ± 0.019	0.714ValTrp: 0.714 ± 0.007	1.604ValTyr: 1.604 ± 0.011	0.0ValXaa: 0.0 ± 0.0
Trp	0.786TrpAla: 0.786 ± 0.007	0.247TrpCys: 0.247 ± 0.004	0.654TrpAsp: 0.654 ± 0.006	0.813TrpGlu: 0.813 ± 0.007	0.441TrpPhe: 0.441 ± 0.005	0.721TrpGly: 0.721 ± 0.008	0.308TrpHis: 0.308 ± 0.004	0.545TrpIle: 0.545 ± 0.006	0.808TrpLys: 0.808 ± 0.007	1.213TrpLeu: 1.213 ± 0.009	0.321TrpMet: 0.321 ± 0.003	0.539TrpAsn: 0.539 ± 0.005	0.525TrpPro: 0.525 ± 0.006	0.535TrpGln: 0.535 ± 0.006	0.723TrpArg: 0.723 ± 0.007	0.87TrpSer: 0.87 ± 0.008	0.663TrpThr: 0.663 ± 0.007	0.69TrpVal: 0.69 ± 0.006	0.198TrpTrp: 0.198 ± 0.004	0.337TrpTyr: 0.337 ± 0.005	0.0TrpXaa: 0.0 ± 0.0
Tyr	1.387TyrAla: 1.387 ± 0.008	0.681TyrCys: 0.681 ± 0.007	1.301TyrAsp: 1.301 ± 0.01	1.735TyrGlu: 1.735 ± 0.012	1.213TyrPhe: 1.213 ± 0.009	1.666TyrGly: 1.666 ± 0.01	0.769TyrHis: 0.769 ± 0.005	1.377TyrIle: 1.377 ± 0.01	1.526TyrLys: 1.526 ± 0.016	2.635TyrLeu: 2.635 ± 0.014	0.606TyrMet: 0.606 ± 0.006	1.118TyrAsn: 1.118 ± 0.009	1.297TyrPro: 1.297 ± 0.01	1.288TyrGln: 1.288 ± 0.009	1.627TyrArg: 1.627 ± 0.011	2.166TyrSer: 2.166 ± 0.011	1.434TyrThr: 1.434 ± 0.01	1.566TyrVal: 1.566 ± 0.01	0.356TyrTrp: 0.356 ± 0.005	0.94TyrTyr: 0.94 ± 0.007	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.0XaaAla: 0.0 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.0XaaAsp: 0.0 ± 0.0	0.0XaaGlu: 0.0 ± 0.0	0.0XaaPhe: 0.0 ± 0.0	0.0XaaGly: 0.0 ± 0.0	0.0XaaHis: 0.0 ± 0.0	0.0XaaIle: 0.0 ± 0.0	0.001XaaLys: 0.001 ± 0.0	0.0XaaLeu: 0.0 ± 0.0	0.0XaaMet: 0.0 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.0XaaPro: 0.0 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.001XaaArg: 0.001 ± 0.0	0.0XaaSer: 0.0 ± 0.0	0.0XaaThr: 0.0 ± 0.0	0.0XaaVal: 0.0 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.115XaaXaa: 0.115 ± 0.039

Statistics based on 40117 proteins (21738302 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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