Amino acid dipepetide frequency for Camelus dromedarius (Dromedary) (Arabian camel)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	7.285AlaAla: 7.285 ± 0.042	1.543AlaCys: 1.543 ± 0.013	2.958AlaAsp: 2.958 ± 0.015	5.02AlaGlu: 5.02 ± 0.024	2.647AlaPhe: 2.647 ± 0.014	5.331AlaGly: 5.331 ± 0.032	1.714AlaHis: 1.714 ± 0.012	2.622AlaIle: 2.622 ± 0.014	3.252AlaLys: 3.252 ± 0.018	7.969AlaLeu: 7.969 ± 0.033	1.503AlaMet: 1.503 ± 0.01	1.86AlaAsn: 1.86 ± 0.012	4.674AlaPro: 4.674 ± 0.031	3.497AlaGln: 3.497 ± 0.021	4.194AlaArg: 4.194 ± 0.023	5.953AlaSer: 5.953 ± 0.026	3.554AlaThr: 3.554 ± 0.018	5.111AlaVal: 5.111 ± 0.022	0.947AlaTrp: 0.947 ± 0.011	1.469AlaTyr: 1.469 ± 0.014	0.001AlaXaa: 0.001 ± 0.0
Cys	1.375CysAla: 1.375 ± 0.011	0.627CysCys: 0.627 ± 0.008	0.997CysAsp: 0.997 ± 0.009	1.231CysGlu: 1.231 ± 0.013	0.883CysPhe: 0.883 ± 0.008	1.684CysGly: 1.684 ± 0.017	0.682CysHis: 0.682 ± 0.008	0.891CysIle: 0.891 ± 0.009	0.986CysLys: 0.986 ± 0.01	2.382CysLeu: 2.382 ± 0.017	0.396CysMet: 0.396 ± 0.005	0.665CysAsn: 0.665 ± 0.008	1.59CysPro: 1.59 ± 0.016	1.076CysGln: 1.076 ± 0.011	1.413CysArg: 1.413 ± 0.012	2.038CysSer: 2.038 ± 0.016	1.074CysThr: 1.074 ± 0.01	1.384CysVal: 1.384 ± 0.012	0.341CysTrp: 0.341 ± 0.005	0.551CysTyr: 0.551 ± 0.007	0.0CysXaa: 0.0 ± 0.0
Asp	2.938AspAla: 2.938 ± 0.017	1.026AspCys: 1.026 ± 0.011	2.497AspAsp: 2.497 ± 0.017	3.3AspGlu: 3.3 ± 0.021	2.058AspPhe: 2.058 ± 0.013	3.264AspGly: 3.264 ± 0.023	1.125AspHis: 1.125 ± 0.01	2.368AspIle: 2.368 ± 0.017	2.284AspLys: 2.284 ± 0.014	5.009AspLeu: 5.009 ± 0.02	1.042AspMet: 1.042 ± 0.01	1.475AspAsn: 1.475 ± 0.011	3.028AspPro: 3.028 ± 0.02	1.816AspGln: 1.816 ± 0.011	2.478AspArg: 2.478 ± 0.015	4.063AspSer: 4.063 ± 0.021	2.419AspThr: 2.419 ± 0.015	3.111AspVal: 3.111 ± 0.021	0.649AspTrp: 0.649 ± 0.007	1.328AspTyr: 1.328 ± 0.011	0.0AspXaa: 0.0 ± 0.0
Glu	5.571GluAla: 5.571 ± 0.026	1.207GluCys: 1.207 ± 0.015	4.363GluAsp: 4.363 ± 0.021	8.001GluGlu: 8.001 ± 0.05	1.949GluPhe: 1.949 ± 0.013	4.527GluGly: 4.527 ± 0.024	1.476GluHis: 1.476 ± 0.011	2.865GluIle: 2.865 ± 0.018	4.955GluLys: 4.955 ± 0.029	6.614GluLeu: 6.614 ± 0.031	1.65GluMet: 1.65 ± 0.011	2.795GluAsn: 2.795 ± 0.018	3.407GluPro: 3.407 ± 0.021	3.188GluGln: 3.188 ± 0.019	4.174GluArg: 4.174 ± 0.026	4.258GluSer: 4.258 ± 0.022	3.285GluThr: 3.285 ± 0.017	4.303GluVal: 4.303 ± 0.022	0.727GluTrp: 0.727 ± 0.007	1.406GluTyr: 1.406 ± 0.013	0.0GluXaa: 0.0 ± 0.0
Phe	1.989PheAla: 1.989 ± 0.012	0.977PheCys: 0.977 ± 0.01	1.546PheAsp: 1.546 ± 0.011	1.926PheGlu: 1.926 ± 0.011	1.645PhePhe: 1.645 ± 0.013	2.159PheGly: 2.159 ± 0.015	1.027PheHis: 1.027 ± 0.009	1.719PheIle: 1.719 ± 0.014	1.622PheLys: 1.622 ± 0.014	4.333PheLeu: 4.333 ± 0.024	0.747PheMet: 0.747 ± 0.008	1.207PheAsn: 1.207 ± 0.009	2.113PhePro: 2.113 ± 0.013	1.854PheGln: 1.854 ± 0.011	1.978PheArg: 1.978 ± 0.014	3.428PheSer: 3.428 ± 0.017	1.958PheThr: 1.958 ± 0.012	2.117PheVal: 2.117 ± 0.013	0.491PheTrp: 0.491 ± 0.006	1.139PheTyr: 1.139 ± 0.01	0.0PheXaa: 0.0 ± 0.0
Gly	5.061GlyAla: 5.061 ± 0.03	1.429GlyCys: 1.429 ± 0.013	3.207GlyAsp: 3.207 ± 0.017	4.171GlyGlu: 4.171 ± 0.023	2.36GlyPhe: 2.36 ± 0.016	5.508GlyGly: 5.508 ± 0.045	1.819GlyHis: 1.819 ± 0.014	2.602GlyIle: 2.602 ± 0.016	3.51GlyLys: 3.51 ± 0.019	6.441GlyLeu: 6.441 ± 0.029	1.263GlyMet: 1.263 ± 0.009	2.159GlyAsn: 2.159 ± 0.015	4.796GlyPro: 4.796 ± 0.04	2.958GlyGln: 2.958 ± 0.016	4.385GlyArg: 4.385 ± 0.028	5.984GlySer: 5.984 ± 0.03	3.667GlyThr: 3.667 ± 0.018	3.799GlyVal: 3.799 ± 0.018	0.953GlyTrp: 0.953 ± 0.01	1.582GlyTyr: 1.582 ± 0.013	0.001GlyXaa: 0.001 ± 0.0
His	1.398HisAla: 1.398 ± 0.01	0.733HisCys: 0.733 ± 0.008	0.857HisAsp: 0.857 ± 0.008	1.234HisGlu: 1.234 ± 0.009	1.128HisPhe: 1.128 ± 0.009	1.599HisGly: 1.599 ± 0.013	0.878HisHis: 0.878 ± 0.009	1.157HisIle: 1.157 ± 0.011	1.141HisLys: 1.141 ± 0.009	3.096HisLeu: 3.096 ± 0.018	0.57HisMet: 0.57 ± 0.007	0.766HisAsn: 0.766 ± 0.007	1.77HisPro: 1.77 ± 0.011	1.293HisGln: 1.293 ± 0.012	1.694HisArg: 1.694 ± 0.012	2.239HisSer: 2.239 ± 0.014	1.397HisThr: 1.397 ± 0.012	1.612HisVal: 1.612 ± 0.01	0.378HisTrp: 0.378 ± 0.005	0.763HisTyr: 0.763 ± 0.007	0.0HisXaa: 0.0 ± 0.0
Ile	2.353IleAla: 2.353 ± 0.015	1.01IleCys: 1.01 ± 0.009	1.818IleAsp: 1.818 ± 0.012	2.301IleGlu: 2.301 ± 0.015	1.78IlePhe: 1.78 ± 0.016	1.985IleGly: 1.985 ± 0.014	1.184IleHis: 1.184 ± 0.01	2.169IleIle: 2.169 ± 0.019	2.29IleLys: 2.29 ± 0.017	4.433IleLeu: 4.433 ± 0.025	0.927IleMet: 0.927 ± 0.008	1.585IleAsn: 1.585 ± 0.012	2.444IlePro: 2.444 ± 0.017	2.15IleGln: 2.15 ± 0.014	2.263IleArg: 2.263 ± 0.014	3.465IleSer: 3.465 ± 0.02	2.445IleThr: 2.445 ± 0.019	2.307IleVal: 2.307 ± 0.017	0.502IleTrp: 0.502 ± 0.007	1.259IleTyr: 1.259 ± 0.01	0.0IleXaa: 0.0 ± 0.0
Lys	3.957LysAla: 3.957 ± 0.021	0.975LysCys: 0.975 ± 0.009	2.846LysAsp: 2.846 ± 0.017	4.794LysGlu: 4.794 ± 0.027	1.601LysPhe: 1.601 ± 0.013	3.106LysGly: 3.106 ± 0.025	1.272LysHis: 1.272 ± 0.011	2.411LysIle: 2.411 ± 0.018	4.217LysLys: 4.217 ± 0.029	4.955LysLeu: 4.955 ± 0.024	1.387LysMet: 1.387 ± 0.012	2.086LysAsn: 2.086 ± 0.015	2.803LysPro: 2.803 ± 0.019	2.416LysGln: 2.416 ± 0.016	3.115LysArg: 3.115 ± 0.017	3.478LysSer: 3.478 ± 0.019	2.81LysThr: 2.81 ± 0.017	3.328LysVal: 3.328 ± 0.02	0.584LysTrp: 0.584 ± 0.007	1.384LysTyr: 1.384 ± 0.012	0.0LysXaa: 0.0 ± 0.0
Leu	7.4LeuAla: 7.4 ± 0.032	2.435LeuCys: 2.435 ± 0.019	4.817LeuAsp: 4.817 ± 0.018	7.436LeuGlu: 7.436 ± 0.037	3.536LeuPhe: 3.536 ± 0.021	6.6LeuGly: 6.6 ± 0.027	2.85LeuHis: 2.85 ± 0.016	3.744LeuIle: 3.744 ± 0.02	5.582LeuLys: 5.582 ± 0.027	11.788LeuLeu: 11.788 ± 0.057	2.035LeuMet: 2.035 ± 0.013	3.306LeuAsn: 3.306 ± 0.021	6.57LeuPro: 6.57 ± 0.028	6.015LeuGln: 6.015 ± 0.03	6.544LeuArg: 6.544 ± 0.03	8.395LeuSer: 8.395 ± 0.032	5.225LeuThr: 5.225 ± 0.019	5.947LeuVal: 5.947 ± 0.028	1.284LeuTrp: 1.284 ± 0.011	2.496LeuTyr: 2.496 ± 0.014	0.0LeuXaa: 0.0 ± 0.0
Met	1.89MetAla: 1.89 ± 0.01	0.402MetCys: 0.402 ± 0.005	1.182MetAsp: 1.182 ± 0.01	1.794MetGlu: 1.794 ± 0.012	0.713MetPhe: 0.713 ± 0.007	1.307MetGly: 1.307 ± 0.01	0.434MetHis: 0.434 ± 0.006	0.794MetIle: 0.794 ± 0.008	1.336MetLys: 1.336 ± 0.01	1.991MetLeu: 1.991 ± 0.014	0.562MetMet: 0.562 ± 0.008	0.797MetAsn: 0.797 ± 0.008	1.08MetPro: 1.08 ± 0.013	0.89MetGln: 0.89 ± 0.009	1.079MetArg: 1.079 ± 0.008	1.47MetSer: 1.47 ± 0.01	1.063MetThr: 1.063 ± 0.009	1.447MetVal: 1.447 ± 0.014	0.247MetTrp: 0.247 ± 0.004	0.538MetTyr: 0.538 ± 0.007	0.0MetXaa: 0.0 ± 0.0
Asn	1.888AsnAla: 1.888 ± 0.013	0.752AsnCys: 0.752 ± 0.007	1.305AsnAsp: 1.305 ± 0.011	1.923AsnGlu: 1.923 ± 0.013	1.361AsnPhe: 1.361 ± 0.011	2.097AsnGly: 2.097 ± 0.015	0.84AsnHis: 0.84 ± 0.008	1.848AsnIle: 1.848 ± 0.014	1.915AsnLys: 1.915 ± 0.013	3.488AsnLeu: 3.488 ± 0.018	0.81AsnMet: 0.81 ± 0.008	1.257AsnAsn: 1.257 ± 0.012	2.062AsnPro: 2.062 ± 0.015	1.494AsnGln: 1.494 ± 0.012	1.708AsnArg: 1.708 ± 0.009	2.72AsnSer: 2.72 ± 0.017	1.726AsnThr: 1.726 ± 0.013	1.998AsnVal: 1.998 ± 0.013	0.416AsnTrp: 0.416 ± 0.005	0.97AsnTyr: 0.97 ± 0.01	0.0AsnXaa: 0.0 ± 0.0
Pro	5.439ProAla: 5.439 ± 0.032	1.275ProCys: 1.275 ± 0.013	2.834ProAsp: 2.834 ± 0.019	4.661ProGlu: 4.661 ± 0.025	1.985ProPhe: 1.985 ± 0.012	5.789ProGly: 5.789 ± 0.056	1.54ProHis: 1.54 ± 0.01	1.721ProIle: 1.721 ± 0.013	2.64ProLys: 2.64 ± 0.023	5.819ProLeu: 5.819 ± 0.026	1.025ProMet: 1.025 ± 0.009	1.622ProAsn: 1.622 ± 0.013	6.271ProPro: 6.271 ± 0.054	2.965ProGln: 2.965 ± 0.02	3.884ProArg: 3.884 ± 0.026	5.839ProSer: 5.839 ± 0.033	3.088ProThr: 3.088 ± 0.02	3.951ProVal: 3.951 ± 0.021	0.819ProTrp: 0.819 ± 0.008	1.328ProTyr: 1.328 ± 0.012	0.002ProXaa: 0.002 ± 0.0
Gln	3.808GlnAla: 3.808 ± 0.022	0.921GlnCys: 0.921 ± 0.009	2.376GlnAsp: 2.376 ± 0.013	4.002GlnGlu: 4.002 ± 0.024	1.314GlnPhe: 1.314 ± 0.01	3.113GlnGly: 3.113 ± 0.018	1.291GlnHis: 1.291 ± 0.01	1.832GlnIle: 1.832 ± 0.013	2.791GlnLys: 2.791 ± 0.018	4.99GlnLeu: 4.99 ± 0.027	1.096GlnMet: 1.096 ± 0.01	1.664GlnAsn: 1.664 ± 0.012	2.832GlnPro: 2.832 ± 0.019	2.915GlnGln: 2.915 ± 0.028	3.081GlnArg: 3.081 ± 0.017	3.019GlnSer: 3.019 ± 0.017	2.233GlnThr: 2.233 ± 0.014	3.02GlnVal: 3.02 ± 0.017	0.559GlnTrp: 0.559 ± 0.006	1.044GlnTyr: 1.044 ± 0.009	0.0GlnXaa: 0.0 ± 0.0
Arg	4.563ArgAla: 4.563 ± 0.026	1.334ArgCys: 1.334 ± 0.013	2.827ArgAsp: 2.827 ± 0.016	4.275ArgGlu: 4.275 ± 0.028	1.864ArgPhe: 1.864 ± 0.012	4.221ArgGly: 4.221 ± 0.028	1.614ArgHis: 1.614 ± 0.011	2.26ArgIle: 2.26 ± 0.012	3.58ArgLys: 3.58 ± 0.017	5.976ArgLeu: 5.976 ± 0.026	1.198ArgMet: 1.198 ± 0.01	1.956ArgAsn: 1.956 ± 0.011	3.674ArgPro: 3.674 ± 0.028	2.791ArgGln: 2.791 ± 0.018	4.813ArgArg: 4.813 ± 0.032	4.302ArgSer: 4.302 ± 0.023	2.865ArgThr: 2.865 ± 0.013	3.492ArgVal: 3.492 ± 0.02	0.784ArgTrp: 0.784 ± 0.008	1.401ArgTyr: 1.401 ± 0.011	0.001ArgXaa: 0.001 ± 0.0
Ser	5.501SerAla: 5.501 ± 0.023	1.879SerCys: 1.879 ± 0.014	3.746SerAsp: 3.746 ± 0.02	5.063SerGlu: 5.063 ± 0.025	3.003SerPhe: 3.003 ± 0.016	5.75SerGly: 5.75 ± 0.028	2.115SerHis: 2.115 ± 0.013	2.949SerIle: 2.949 ± 0.014	3.732SerLys: 3.732 ± 0.02	8.61SerLeu: 8.61 ± 0.034	1.501SerMet: 1.501 ± 0.01	2.316SerAsn: 2.316 ± 0.015	6.15SerPro: 6.15 ± 0.036	3.833SerGln: 3.833 ± 0.02	4.718SerArg: 4.718 ± 0.025	8.779SerSer: 8.779 ± 0.045	4.156SerThr: 4.156 ± 0.022	4.947SerVal: 4.947 ± 0.018	1.182SerTrp: 1.182 ± 0.01	1.921SerTyr: 1.921 ± 0.013	0.001SerXaa: 0.001 ± 0.0
Thr	3.844ThrAla: 3.844 ± 0.019	1.302ThrCys: 1.302 ± 0.015	2.334ThrAsp: 2.334 ± 0.014	3.403ThrGlu: 3.403 ± 0.018	2.06ThrPhe: 2.06 ± 0.011	3.481ThrGly: 3.481 ± 0.02	1.261ThrHis: 1.261 ± 0.01	2.225ThrIle: 2.225 ± 0.016	2.391ThrLys: 2.391 ± 0.016	5.389ThrLeu: 5.389 ± 0.022	1.046ThrMet: 1.046 ± 0.007	1.499ThrAsn: 1.499 ± 0.013	3.541ThrPro: 3.541 ± 0.021	2.258ThrGln: 2.258 ± 0.014	2.568ThrArg: 2.568 ± 0.013	4.368ThrSer: 4.368 ± 0.02	2.811ThrThr: 2.811 ± 0.02	3.794ThrVal: 3.794 ± 0.021	0.75ThrTrp: 0.75 ± 0.008	1.319ThrTyr: 1.319 ± 0.009	0.001ThrXaa: 0.001 ± 0.0
Val	4.554ValAla: 4.554 ± 0.021	1.532ValCys: 1.532 ± 0.013	2.943ValAsp: 2.943 ± 0.02	3.812ValGlu: 3.812 ± 0.016	2.462ValPhe: 2.462 ± 0.015	3.655ValGly: 3.655 ± 0.019	1.561ValHis: 1.561 ± 0.009	2.775ValIle: 2.775 ± 0.014	3.224ValLys: 3.224 ± 0.019	6.768ValLeu: 6.768 ± 0.028	1.388ValMet: 1.388 ± 0.013	2.105ValAsn: 2.105 ± 0.013	3.86ValPro: 3.86 ± 0.02	2.779ValGln: 2.779 ± 0.014	3.354ValArg: 3.354 ± 0.014	5.102ValSer: 5.102 ± 0.022	3.79ValThr: 3.79 ± 0.021	4.211ValVal: 4.211 ± 0.024	0.778ValTrp: 0.778 ± 0.008	1.565ValTyr: 1.565 ± 0.01	0.001ValXaa: 0.001 ± 0.0
Trp	0.96TrpAla: 0.96 ± 0.009	0.264TrpCys: 0.264 ± 0.004	0.693TrpAsp: 0.693 ± 0.007	0.877TrpGlu: 0.877 ± 0.008	0.453TrpPhe: 0.453 ± 0.005	0.938TrpGly: 0.938 ± 0.012	0.32TrpHis: 0.32 ± 0.005	0.516TrpIle: 0.516 ± 0.007	0.808TrpLys: 0.808 ± 0.007	1.323TrpLeu: 1.323 ± 0.011	0.316TrpMet: 0.316 ± 0.004	0.504TrpAsn: 0.504 ± 0.007	0.631TrpPro: 0.631 ± 0.006	0.56TrpGln: 0.56 ± 0.006	0.839TrpArg: 0.839 ± 0.009	0.925TrpSer: 0.925 ± 0.009	0.703TrpThr: 0.703 ± 0.009	0.793TrpVal: 0.793 ± 0.009	0.209TrpTrp: 0.209 ± 0.004	0.311TrpTyr: 0.311 ± 0.004	0.0TrpXaa: 0.0 ± 0.0
Tyr	1.33TyrAla: 1.33 ± 0.01	0.643TyrCys: 0.643 ± 0.007	1.134TyrAsp: 1.134 ± 0.01	1.495TyrGlu: 1.495 ± 0.011	1.172TyrPhe: 1.172 ± 0.011	1.541TyrGly: 1.541 ± 0.013	0.701TyrHis: 0.701 ± 0.007	1.202TyrIle: 1.202 ± 0.01	1.266TyrLys: 1.266 ± 0.014	2.632TyrLeu: 2.632 ± 0.016	0.554TyrMet: 0.554 ± 0.006	0.955TyrAsn: 0.955 ± 0.009	1.201TyrPro: 1.201 ± 0.01	1.152TyrGln: 1.152 ± 0.009	1.505TyrArg: 1.505 ± 0.013	2.028TyrSer: 2.028 ± 0.012	1.376TyrThr: 1.376 ± 0.01	1.546TyrVal: 1.546 ± 0.012	0.345TyrTrp: 0.345 ± 0.006	0.83TyrTyr: 0.83 ± 0.01	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.002XaaAla: 0.002 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.0XaaAsp: 0.0 ± 0.0	0.0XaaGlu: 0.0 ± 0.0	0.0XaaPhe: 0.0 ± 0.0	0.001XaaGly: 0.001 ± 0.0	0.0XaaHis: 0.0 ± 0.0	0.0XaaIle: 0.0 ± 0.0	0.001XaaLys: 0.001 ± 0.0	0.001XaaLeu: 0.001 ± 0.0	0.0XaaMet: 0.0 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.001XaaPro: 0.001 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.001XaaArg: 0.001 ± 0.0	0.0XaaSer: 0.0 ± 0.0	0.0XaaThr: 0.0 ± 0.0	0.0XaaVal: 0.0 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.036XaaXaa: 0.036 ± 0.005

Statistics based on 31080 proteins (16143790 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

---