Amino acid dipepetide frequency for Beet yellows virus (isolate Ukraine) (BYV) (Sugar beet yellows virus)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
4.283AlaAla: 4.283 ± 1.076
1.557AlaCys: 1.557 ± 0.331
1.817AlaAsp: 1.817 ± 0.516
3.245AlaGlu: 3.245 ± 0.913
4.153AlaPhe: 4.153 ± 0.428
2.985AlaGly: 2.985 ± 0.526
0.519AlaHis: 0.519 ± 0.274
3.245AlaIle: 3.245 ± 0.561
3.634AlaLys: 3.634 ± 1.01
8.177AlaLeu: 8.177 ± 0.781
0.909AlaMet: 0.909 ± 0.339
1.947AlaAsn: 1.947 ± 0.189
2.466AlaPro: 2.466 ± 0.395
2.855AlaGln: 2.855 ± 0.327
4.283AlaArg: 4.283 ± 0.486
6.749AlaSer: 6.749 ± 0.765
2.596AlaThr: 2.596 ± 1.045
4.543AlaVal: 4.543 ± 0.867
0.649AlaTrp: 0.649 ± 0.232
3.504AlaTyr: 3.504 ± 0.57
0.0AlaXaa: 0.0 ± 0.0
Cys
2.077CysAla: 2.077 ± 0.46
0.909CysCys: 0.909 ± 0.207
2.077CysAsp: 2.077 ± 0.36
1.038CysGlu: 1.038 ± 0.233
1.947CysPhe: 1.947 ± 0.355
1.038CysGly: 1.038 ± 0.491
0.26CysHis: 0.26 ± 0.087
0.389CysIle: 0.389 ± 0.145
1.038CysLys: 1.038 ± 0.267
2.336CysLeu: 2.336 ± 1.165
0.26CysMet: 0.26 ± 0.408
1.168CysAsn: 1.168 ± 0.28
1.038CysPro: 1.038 ± 0.347
0.13CysGln: 0.13 ± 0.221
1.168CysArg: 1.168 ± 0.322
2.596CysSer: 2.596 ± 0.527
0.649CysThr: 0.649 ± 0.225
3.115CysVal: 3.115 ± 0.522
0.389CysTrp: 0.389 ± 0.131
0.779CysTyr: 0.779 ± 0.164
0.0CysXaa: 0.0 ± 0.0
Asp
3.634AspAla: 3.634 ± 0.584
1.168AspCys: 1.168 ± 0.513
2.596AspAsp: 2.596 ± 0.534
5.321AspGlu: 5.321 ± 0.657
4.153AspPhe: 4.153 ± 0.835
1.687AspGly: 1.687 ± 0.197
1.298AspHis: 1.298 ± 0.288
1.557AspIle: 1.557 ± 0.331
1.557AspLys: 1.557 ± 0.611
4.153AspLeu: 4.153 ± 0.591
1.298AspMet: 1.298 ± 0.168
1.817AspAsn: 1.817 ± 0.32
0.649AspPro: 0.649 ± 0.22
0.389AspGln: 0.389 ± 0.217
1.947AspArg: 1.947 ± 0.554
4.672AspSer: 4.672 ± 0.72
1.687AspThr: 1.687 ± 0.229
4.543AspVal: 4.543 ± 0.589
0.26AspTrp: 0.26 ± 0.087
2.336AspTyr: 2.336 ± 0.674
0.0AspXaa: 0.0 ± 0.0
Glu
5.321GluAla: 5.321 ± 0.622
1.817GluCys: 1.817 ± 0.276
3.245GluAsp: 3.245 ± 0.827
5.191GluGlu: 5.191 ± 0.495
4.023GluPhe: 4.023 ± 0.897
1.038GluGly: 1.038 ± 0.22
1.298GluHis: 1.298 ± 0.199
4.283GluIle: 4.283 ± 0.71
3.374GluLys: 3.374 ± 0.302
6.1GluLeu: 6.1 ± 2.625
1.557GluMet: 1.557 ± 0.315
2.985GluAsn: 2.985 ± 0.708
1.947GluPro: 1.947 ± 0.27
1.168GluGln: 1.168 ± 0.282
4.413GluArg: 4.413 ± 0.722
5.97GluSer: 5.97 ± 0.41
3.764GluThr: 3.764 ± 0.657
5.97GluVal: 5.97 ± 0.406
0.779GluTrp: 0.779 ± 0.197
3.374GluTyr: 3.374 ± 0.537
0.0GluXaa: 0.0 ± 0.0
Phe
2.077PheAla: 2.077 ± 0.582
1.687PheCys: 1.687 ± 0.289
2.726PheAsp: 2.726 ± 0.501
4.153PheGlu: 4.153 ± 1.24
4.153PhePhe: 4.153 ± 0.645
4.283PheGly: 4.283 ± 0.773
1.687PheHis: 1.687 ± 0.336
2.077PheIle: 2.077 ± 0.458
3.245PheLys: 3.245 ± 0.424
8.566PheLeu: 8.566 ± 0.793
1.168PheMet: 1.168 ± 0.313
1.557PheAsn: 1.557 ± 0.611
2.077PhePro: 2.077 ± 0.273
0.649PheGln: 0.649 ± 0.375
4.543PheArg: 4.543 ± 0.937
6.619PheSer: 6.619 ± 0.456
2.726PheThr: 2.726 ± 0.363
6.749PheVal: 6.749 ± 0.54
0.0PheTrp: 0.0 ± 0.0
1.947PheTyr: 1.947 ± 0.328
0.0PheXaa: 0.0 ± 0.0
Gly
2.206GlyAla: 2.206 ± 0.451
1.168GlyCys: 1.168 ± 0.308
3.245GlyAsp: 3.245 ± 0.577
5.191GlyGlu: 5.191 ± 0.744
2.726GlyPhe: 2.726 ± 0.308
5.062GlyGly: 5.062 ± 1.2
0.389GlyHis: 0.389 ± 0.145
1.038GlyIle: 1.038 ± 0.323
2.985GlyLys: 2.985 ± 1.243
5.84GlyLeu: 5.84 ± 0.691
0.649GlyMet: 0.649 ± 0.361
1.817GlyAsn: 1.817 ± 0.272
1.038GlyPro: 1.038 ± 0.215
0.909GlyGln: 0.909 ± 0.248
2.466GlyArg: 2.466 ± 0.401
5.062GlySer: 5.062 ± 0.641
2.466GlyThr: 2.466 ± 0.984
4.672GlyVal: 4.672 ± 0.785
0.26GlyTrp: 0.26 ± 0.167
1.038GlyTyr: 1.038 ± 0.362
0.0GlyXaa: 0.0 ± 0.0
His
2.077HisAla: 2.077 ± 0.379
0.909HisCys: 0.909 ± 0.517
0.649HisAsp: 0.649 ± 0.259
1.038HisGlu: 1.038 ± 0.262
0.909HisPhe: 0.909 ± 0.272
1.168HisGly: 1.168 ± 0.652
0.389HisHis: 0.389 ± 0.177
1.038HisIle: 1.038 ± 0.347
1.557HisLys: 1.557 ± 0.297
1.038HisLeu: 1.038 ± 0.266
0.13HisMet: 0.13 ± 0.156
0.389HisAsn: 0.389 ± 0.51
1.557HisPro: 1.557 ± 0.414
0.0HisGln: 0.0 ± 0.0
0.649HisArg: 0.649 ± 0.209
1.557HisSer: 1.557 ± 0.341
0.909HisThr: 0.909 ± 0.228
1.428HisVal: 1.428 ± 0.321
0.0HisTrp: 0.0 ± 0.0
0.389HisTyr: 0.389 ± 0.363
0.0HisXaa: 0.0 ± 0.0
Ile
2.726IleAla: 2.726 ± 0.568
0.519IleCys: 0.519 ± 0.279
2.466IleAsp: 2.466 ± 0.273
3.245IleGlu: 3.245 ± 0.688
1.817IlePhe: 1.817 ± 0.311
1.947IleGly: 1.947 ± 0.531
1.428IleHis: 1.428 ± 0.242
1.687IleIle: 1.687 ± 0.395
2.596IleLys: 2.596 ± 0.187
4.283IleLeu: 4.283 ± 0.521
0.779IleMet: 0.779 ± 0.214
1.817IleAsn: 1.817 ± 0.555
1.817IlePro: 1.817 ± 0.662
0.13IleGln: 0.13 ± 0.163
3.245IleArg: 3.245 ± 0.663
2.855IleSer: 2.855 ± 1.429
3.245IleThr: 3.245 ± 0.663
2.855IleVal: 2.855 ± 0.599
0.13IleTrp: 0.13 ± 0.191
0.519IleTyr: 0.519 ± 0.173
0.0IleXaa: 0.0 ± 0.0
Lys
3.245LysAla: 3.245 ± 0.715
1.038LysCys: 1.038 ± 0.314
3.115LysAsp: 3.115 ± 0.439
3.115LysGlu: 3.115 ± 0.777
4.153LysPhe: 4.153 ± 0.685
1.947LysGly: 1.947 ± 0.36
1.947LysHis: 1.947 ± 0.257
2.336LysIle: 2.336 ± 0.416
2.855LysLys: 2.855 ± 0.714
6.36LysLeu: 6.36 ± 1.72
0.519LysMet: 0.519 ± 0.221
2.077LysAsn: 2.077 ± 0.778
2.855LysPro: 2.855 ± 0.475
0.909LysGln: 0.909 ± 0.315
5.321LysArg: 5.321 ± 0.395
5.191LysSer: 5.191 ± 0.73
4.283LysThr: 4.283 ± 0.72
6.1LysVal: 6.1 ± 0.34
0.0LysTrp: 0.0 ± 0.0
1.038LysTyr: 1.038 ± 0.342
0.0LysXaa: 0.0 ± 0.0
Leu
6.36LeuAla: 6.36 ± 0.804
2.726LeuCys: 2.726 ± 0.595
4.802LeuAsp: 4.802 ± 0.601
4.672LeuGlu: 4.672 ± 0.382
5.191LeuPhe: 5.191 ± 0.677
4.543LeuGly: 4.543 ± 1.483
1.168LeuHis: 1.168 ± 0.577
3.894LeuIle: 3.894 ± 0.768
7.008LeuLys: 7.008 ± 1.534
9.994LeuLeu: 9.994 ± 1.186
1.038LeuMet: 1.038 ± 0.2
5.97LeuAsn: 5.97 ± 1.273
5.321LeuPro: 5.321 ± 0.487
3.504LeuGln: 3.504 ± 0.395
6.1LeuArg: 6.1 ± 0.942
12.589LeuSer: 12.589 ± 1.328
7.398LeuThr: 7.398 ± 0.381
6.1LeuVal: 6.1 ± 0.507
0.26LeuTrp: 0.26 ± 0.327
3.504LeuTyr: 3.504 ± 0.795
0.0LeuXaa: 0.0 ± 0.0
Met
1.038MetAla: 1.038 ± 0.269
0.649MetCys: 0.649 ± 0.28
0.13MetAsp: 0.13 ± 0.191
0.649MetGlu: 0.649 ± 0.33
0.909MetPhe: 0.909 ± 0.264
1.428MetGly: 1.428 ± 0.342
0.26MetHis: 0.26 ± 0.272
0.0MetIle: 0.0 ± 0.0
1.298MetLys: 1.298 ± 0.296
1.687MetLeu: 1.687 ± 0.251
0.0MetMet: 0.0 ± 0.0
0.649MetAsn: 0.649 ± 0.349
0.389MetPro: 0.389 ± 0.131
0.389MetGln: 0.389 ± 0.145
0.26MetArg: 0.26 ± 0.337
1.038MetSer: 1.038 ± 0.335
0.779MetThr: 0.779 ± 0.563
1.947MetVal: 1.947 ± 0.481
0.0MetTrp: 0.0 ± 0.0
0.909MetTyr: 0.909 ± 0.258
0.0MetXaa: 0.0 ± 0.0
Asn
2.726AsnAla: 2.726 ± 0.479
1.168AsnCys: 1.168 ± 0.187
0.519AsnAsp: 0.519 ± 0.571
2.855AsnGlu: 2.855 ± 0.337
3.115AsnPhe: 3.115 ± 0.658
1.168AsnGly: 1.168 ± 0.326
0.779AsnHis: 0.779 ± 0.291
1.687AsnIle: 1.687 ± 0.854
2.077AsnLys: 2.077 ± 0.853
4.672AsnLeu: 4.672 ± 0.385
1.038AsnMet: 1.038 ± 0.364
0.519AsnAsn: 0.519 ± 0.221
1.557AsnPro: 1.557 ± 0.755
0.519AsnGln: 0.519 ± 0.184
3.245AsnArg: 3.245 ± 0.613
4.932AsnSer: 4.932 ± 0.405
0.779AsnThr: 0.779 ± 0.31
3.374AsnVal: 3.374 ± 0.924
0.13AsnTrp: 0.13 ± 0.163
1.817AsnTyr: 1.817 ± 0.464
0.0AsnXaa: 0.0 ± 0.0
Pro
2.726ProAla: 2.726 ± 0.329
1.038ProCys: 1.038 ± 0.215
0.649ProAsp: 0.649 ± 0.208
2.206ProGlu: 2.206 ± 0.38
2.726ProPhe: 2.726 ± 0.564
3.245ProGly: 3.245 ± 0.587
1.038ProHis: 1.038 ± 0.142
2.466ProIle: 2.466 ± 0.359
2.596ProLys: 2.596 ± 0.349
4.932ProLeu: 4.932 ± 0.727
0.26ProMet: 0.26 ± 0.337
0.909ProAsn: 0.909 ± 0.65
1.557ProPro: 1.557 ± 0.303
1.298ProGln: 1.298 ± 0.276
3.504ProArg: 3.504 ± 1.037
3.374ProSer: 3.374 ± 0.349
2.855ProThr: 2.855 ± 0.324
2.596ProVal: 2.596 ± 0.375
0.519ProTrp: 0.519 ± 0.173
0.779ProTyr: 0.779 ± 0.29
0.0ProXaa: 0.0 ± 0.0
Gln
1.557GlnAla: 1.557 ± 0.352
0.389GlnCys: 0.389 ± 0.17
0.26GlnAsp: 0.26 ± 0.167
1.557GlnGlu: 1.557 ± 0.253
0.519GlnPhe: 0.519 ± 0.173
1.168GlnGly: 1.168 ± 0.156
0.389GlnHis: 0.389 ± 0.177
1.298GlnIle: 1.298 ± 0.373
0.779GlnLys: 0.779 ± 0.508
1.557GlnLeu: 1.557 ± 0.424
0.0GlnMet: 0.0 ± 0.0
0.519GlnAsn: 0.519 ± 0.176
1.038GlnPro: 1.038 ± 0.22
0.519GlnGln: 0.519 ± 0.329
1.817GlnArg: 1.817 ± 0.677
1.557GlnSer: 1.557 ± 0.561
1.168GlnThr: 1.168 ± 0.359
1.298GlnVal: 1.298 ± 0.434
0.0GlnTrp: 0.0 ± 0.0
1.298GlnTyr: 1.298 ± 0.434
0.0GlnXaa: 0.0 ± 0.0
Arg
3.374ArgAla: 3.374 ± 0.608
1.038ArgCys: 1.038 ± 0.338
2.466ArgAsp: 2.466 ± 0.636
4.672ArgGlu: 4.672 ± 0.55
4.802ArgPhe: 4.802 ± 0.993
4.153ArgGly: 4.153 ± 0.481
0.909ArgHis: 0.909 ± 0.261
1.687ArgIle: 1.687 ± 0.647
4.672ArgLys: 4.672 ± 0.397
6.879ArgLeu: 6.879 ± 0.527
0.909ArgMet: 0.909 ± 0.227
2.466ArgAsn: 2.466 ± 0.633
3.374ArgPro: 3.374 ± 0.882
1.298ArgGln: 1.298 ± 0.434
5.84ArgArg: 5.84 ± 1.07
6.619ArgSer: 6.619 ± 1.257
4.543ArgThr: 4.543 ± 0.442
7.398ArgVal: 7.398 ± 0.965
0.389ArgTrp: 0.389 ± 0.413
1.038ArgTyr: 1.038 ± 0.267
0.0ArgXaa: 0.0 ± 0.0
Ser
8.825SerAla: 8.825 ± 1.472
2.206SerCys: 2.206 ± 0.238
5.84SerAsp: 5.84 ± 0.687
7.138SerGlu: 7.138 ± 0.786
4.413SerPhe: 4.413 ± 0.623
6.489SerGly: 6.489 ± 0.828
1.038SerHis: 1.038 ± 0.267
4.153SerIle: 4.153 ± 0.633
5.581SerLys: 5.581 ± 0.891
7.138SerLeu: 7.138 ± 1.17
1.298SerMet: 1.298 ± 0.183
4.153SerAsn: 4.153 ± 0.455
4.153SerPro: 4.153 ± 0.44
1.428SerGln: 1.428 ± 0.473
8.955SerArg: 8.955 ± 1.424
12.2SerSer: 12.2 ± 0.973
4.672SerThr: 4.672 ± 0.64
10.513SerVal: 10.513 ± 1.143
0.519SerTrp: 0.519 ± 0.173
3.634SerTyr: 3.634 ± 0.612
0.0SerXaa: 0.0 ± 0.0
Thr
2.726ThrAla: 2.726 ± 0.327
1.168ThrCys: 1.168 ± 0.231
1.687ThrAsp: 1.687 ± 0.458
3.634ThrGlu: 3.634 ± 0.571
6.619ThrPhe: 6.619 ± 1.209
1.687ThrGly: 1.687 ± 1.146
0.909ThrHis: 0.909 ± 0.43
1.947ThrIle: 1.947 ± 0.621
3.634ThrLys: 3.634 ± 0.808
4.802ThrLeu: 4.802 ± 0.77
0.779ThrMet: 0.779 ± 0.26
2.336ThrAsn: 2.336 ± 0.667
2.726ThrPro: 2.726 ± 0.65
1.038ThrGln: 1.038 ± 0.142
2.596ThrArg: 2.596 ± 0.701
4.543ThrSer: 4.543 ± 0.886
2.726ThrThr: 2.726 ± 0.45
3.764ThrVal: 3.764 ± 0.63
0.26ThrTrp: 0.26 ± 0.186
2.077ThrTyr: 2.077 ± 0.263
0.0ThrXaa: 0.0 ± 0.0
Val
6.489ValAla: 6.489 ± 0.636
1.687ValCys: 1.687 ± 0.28
5.191ValAsp: 5.191 ± 0.308
5.062ValGlu: 5.062 ± 0.597
4.543ValPhe: 4.543 ± 0.557
3.504ValGly: 3.504 ± 1.003
1.557ValHis: 1.557 ± 0.52
3.894ValIle: 3.894 ± 0.568
6.23ValLys: 6.23 ± 1.327
8.047ValLeu: 8.047 ± 1.028
0.779ValMet: 0.779 ± 0.241
4.283ValAsn: 4.283 ± 0.643
4.413ValPro: 4.413 ± 0.469
1.298ValGln: 1.298 ± 0.343
5.581ValArg: 5.581 ± 0.532
10.772ValSer: 10.772 ± 1.119
2.985ValThr: 2.985 ± 0.591
8.177ValVal: 8.177 ± 1.483
0.649ValTrp: 0.649 ± 0.22
2.726ValTyr: 2.726 ± 0.61
0.0ValXaa: 0.0 ± 0.0
Trp
0.0TrpAla: 0.0 ± 0.0
0.26TrpCys: 0.26 ± 0.087
0.26TrpAsp: 0.26 ± 0.087
0.389TrpGlu: 0.389 ± 0.131
0.519TrpPhe: 0.519 ± 0.29
0.389TrpGly: 0.389 ± 0.131
0.0TrpHis: 0.0 ± 0.0
0.26TrpIle: 0.26 ± 0.214
0.389TrpLys: 0.389 ± 0.49
0.519TrpLeu: 0.519 ± 0.173
0.519TrpMet: 0.519 ± 0.299
0.0TrpAsn: 0.0 ± 0.0
0.0TrpPro: 0.0 ± 0.0
0.0TrpGln: 0.0 ± 0.0
0.0TrpArg: 0.0 ± 0.0
0.649TrpSer: 0.649 ± 0.15
0.0TrpThr: 0.0 ± 0.0
1.428TrpVal: 1.428 ± 0.311
0.0TrpTrp: 0.0 ± 0.0
0.0TrpTyr: 0.0 ± 0.0
0.0TrpXaa: 0.0 ± 0.0
Tyr
0.649TyrAla: 0.649 ± 0.22
1.298TyrCys: 1.298 ± 0.418
2.985TyrAsp: 2.985 ± 0.525
3.245TyrGlu: 3.245 ± 0.514
1.298TyrPhe: 1.298 ± 0.314
1.298TyrGly: 1.298 ± 0.279
0.519TyrHis: 0.519 ± 0.173
1.168TyrIle: 1.168 ± 0.275
1.298TyrLys: 1.298 ± 0.464
5.062TyrLeu: 5.062 ± 0.987
0.389TyrMet: 0.389 ± 0.17
1.298TyrAsn: 1.298 ± 0.417
1.298TyrPro: 1.298 ± 0.276
0.26TyrGln: 0.26 ± 0.441
2.596TyrArg: 2.596 ± 0.411
4.802TyrSer: 4.802 ± 0.424
1.428TyrThr: 1.428 ± 0.354
1.428TyrVal: 1.428 ± 0.59
0.389TyrTrp: 0.389 ± 0.131
2.077TyrTyr: 2.077 ± 0.466
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 9 proteins (7706 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski