Amino acid dipepetide frequency for Tursiops truncatus papillomavirus 5

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
3.528AlaAla: 3.528 ± 1.114
1.96AlaCys: 1.96 ± 0.824
3.92AlaAsp: 3.92 ± 1.298
1.96AlaGlu: 1.96 ± 0.909
2.352AlaPhe: 2.352 ± 0.401
4.312AlaGly: 4.312 ± 0.764
1.176AlaHis: 1.176 ± 0.576
1.96AlaIle: 1.96 ± 0.569
3.136AlaLys: 3.136 ± 0.78
7.056AlaLeu: 7.056 ± 1.435
1.96AlaMet: 1.96 ± 0.396
3.92AlaAsn: 3.92 ± 1.319
3.92AlaPro: 3.92 ± 1.39
1.96AlaGln: 1.96 ± 0.622
3.136AlaArg: 3.136 ± 0.874
2.744AlaSer: 2.744 ± 0.616
3.136AlaThr: 3.136 ± 1.515
2.352AlaVal: 2.352 ± 0.608
0.0AlaTrp: 0.0 ± 0.0
1.176AlaTyr: 1.176 ± 0.612
0.0AlaXaa: 0.0 ± 0.0
Cys
0.784CysAla: 0.784 ± 0.651
0.392CysCys: 0.392 ± 0.325
2.744CysAsp: 2.744 ± 1.044
1.176CysGlu: 1.176 ± 0.617
1.176CysPhe: 1.176 ± 0.976
0.784CysGly: 0.784 ± 0.42
0.392CysHis: 0.392 ± 0.533
1.176CysIle: 1.176 ± 0.584
1.176CysLys: 1.176 ± 0.661
2.744CysLeu: 2.744 ± 1.169
0.392CysMet: 0.392 ± 0.428
1.176CysAsn: 1.176 ± 1.014
1.568CysPro: 1.568 ± 0.816
0.392CysGln: 0.392 ± 0.325
1.96CysArg: 1.96 ± 0.58
1.96CysSer: 1.96 ± 1.004
3.136CysThr: 3.136 ± 1.018
2.352CysVal: 2.352 ± 1.67
1.568CysTrp: 1.568 ± 0.878
0.0CysTyr: 0.0 ± 0.0
0.0CysXaa: 0.0 ± 0.0
Asp
3.528AspAla: 3.528 ± 0.882
3.528AspCys: 3.528 ± 1.322
3.136AspAsp: 3.136 ± 1.277
3.136AspGlu: 3.136 ± 2.232
2.352AspPhe: 2.352 ± 0.654
4.312AspGly: 4.312 ± 1.508
0.392AspHis: 0.392 ± 0.643
3.92AspIle: 3.92 ± 1.381
3.136AspLys: 3.136 ± 1.309
6.272AspLeu: 6.272 ± 2.115
1.568AspMet: 1.568 ± 0.927
2.744AspAsn: 2.744 ± 0.741
5.488AspPro: 5.488 ± 1.718
1.176AspGln: 1.176 ± 0.382
1.176AspArg: 1.176 ± 0.5
3.92AspSer: 3.92 ± 0.905
7.056AspThr: 7.056 ± 1.552
3.92AspVal: 3.92 ± 2.291
1.176AspTrp: 1.176 ± 0.721
1.568AspTyr: 1.568 ± 1.301
0.0AspXaa: 0.0 ± 0.0
Glu
1.96GluAla: 1.96 ± 1.181
0.784GluCys: 0.784 ± 0.651
5.88GluAsp: 5.88 ± 2.543
3.92GluGlu: 3.92 ± 0.992
1.96GluPhe: 1.96 ± 0.577
5.096GluGly: 5.096 ± 1.268
1.568GluHis: 1.568 ± 0.609
1.176GluIle: 1.176 ± 0.576
4.704GluLys: 4.704 ± 1.431
4.312GluLeu: 4.312 ± 1.28
1.96GluMet: 1.96 ± 0.59
2.744GluAsn: 2.744 ± 1.157
1.568GluPro: 1.568 ± 0.722
2.352GluGln: 2.352 ± 1.378
1.96GluArg: 1.96 ± 0.706
2.352GluSer: 2.352 ± 1.036
6.272GluThr: 6.272 ± 1.247
2.744GluVal: 2.744 ± 0.855
0.0GluTrp: 0.0 ± 0.0
1.176GluTyr: 1.176 ± 0.434
0.0GluXaa: 0.0 ± 0.0
Phe
1.96PheAla: 1.96 ± 0.889
0.392PheCys: 0.392 ± 0.452
3.136PheAsp: 3.136 ± 1.415
1.96PheGlu: 1.96 ± 0.787
1.96PhePhe: 1.96 ± 0.577
2.744PheGly: 2.744 ± 0.488
1.176PheHis: 1.176 ± 0.584
2.744PheIle: 2.744 ± 1.083
3.528PheLys: 3.528 ± 1.286
6.664PheLeu: 6.664 ± 1.317
0.392PheMet: 0.392 ± 0.399
1.568PheAsn: 1.568 ± 0.931
1.568PhePro: 1.568 ± 0.682
2.744PheGln: 2.744 ± 0.936
0.392PheArg: 0.392 ± 0.533
3.136PheSer: 3.136 ± 0.88
1.568PheThr: 1.568 ± 0.62
2.744PheVal: 2.744 ± 1.124
0.784PheTrp: 0.784 ± 0.651
2.352PheTyr: 2.352 ± 1.171
0.0PheXaa: 0.0 ± 0.0
Gly
3.92GlyAla: 3.92 ± 0.888
1.96GlyCys: 1.96 ± 0.799
5.096GlyAsp: 5.096 ± 0.989
4.704GlyGlu: 4.704 ± 1.324
1.96GlyPhe: 1.96 ± 0.934
8.624GlyGly: 8.624 ± 2.021
1.568GlyHis: 1.568 ± 0.873
3.136GlyIle: 3.136 ± 0.762
4.312GlyLys: 4.312 ± 1.557
3.528GlyLeu: 3.528 ± 0.921
0.0GlyMet: 0.0 ± 0.0
5.488GlyAsn: 5.488 ± 0.778
3.528GlyPro: 3.528 ± 1.273
3.528GlyGln: 3.528 ± 1.262
3.92GlyArg: 3.92 ± 1.006
4.704GlySer: 4.704 ± 1.487
4.312GlyThr: 4.312 ± 1.079
1.568GlyVal: 1.568 ± 0.687
0.0GlyTrp: 0.0 ± 0.0
1.568GlyTyr: 1.568 ± 0.636
0.0GlyXaa: 0.0 ± 0.0
His
0.784HisAla: 0.784 ± 0.408
0.0HisCys: 0.0 ± 0.0
0.784HisAsp: 0.784 ± 0.39
0.392HisGlu: 0.392 ± 0.302
1.176HisPhe: 1.176 ± 0.513
1.176HisGly: 1.176 ± 0.576
3.136HisHis: 3.136 ± 5.144
1.96HisIle: 1.96 ± 0.59
1.176HisLys: 1.176 ± 0.783
3.136HisLeu: 3.136 ± 1.024
1.176HisMet: 1.176 ± 0.956
0.392HisAsn: 0.392 ± 0.324
3.528HisPro: 3.528 ± 2.086
1.568HisGln: 1.568 ± 2.572
1.96HisArg: 1.96 ± 0.707
1.568HisSer: 1.568 ± 0.63
0.784HisThr: 0.784 ± 0.605
1.176HisVal: 1.176 ± 0.712
0.784HisTrp: 0.784 ± 0.444
0.784HisTyr: 0.784 ± 0.601
0.0HisXaa: 0.0 ± 0.0
Ile
2.352IleAla: 2.352 ± 0.462
1.568IleCys: 1.568 ± 0.673
2.352IleAsp: 2.352 ± 0.813
2.744IleGlu: 2.744 ± 0.877
2.352IlePhe: 2.352 ± 0.653
3.528IleGly: 3.528 ± 1.368
0.392IleHis: 0.392 ± 0.324
1.176IleIle: 1.176 ± 0.714
0.392IleLys: 0.392 ± 0.302
5.096IleLeu: 5.096 ± 1.342
1.568IleMet: 1.568 ± 0.807
1.568IleAsn: 1.568 ± 0.433
2.352IlePro: 2.352 ± 0.861
2.744IleGln: 2.744 ± 0.936
2.744IleArg: 2.744 ± 0.816
3.92IleSer: 3.92 ± 1.176
3.528IleThr: 3.528 ± 0.93
1.96IleVal: 1.96 ± 0.698
0.784IleTrp: 0.784 ± 0.752
1.96IleTyr: 1.96 ± 0.87
0.0IleXaa: 0.0 ± 0.0
Lys
3.92LysAla: 3.92 ± 1.138
3.136LysCys: 3.136 ± 1.649
3.136LysAsp: 3.136 ± 1.058
1.96LysGlu: 1.96 ± 1.275
2.744LysPhe: 2.744 ± 0.69
0.784LysGly: 0.784 ± 0.42
1.96LysHis: 1.96 ± 0.67
3.528LysIle: 3.528 ± 0.971
4.312LysLys: 4.312 ± 1.597
1.176LysLeu: 1.176 ± 0.661
0.0LysMet: 0.0 ± 0.0
3.136LysAsn: 3.136 ± 1.325
2.352LysPro: 2.352 ± 0.642
3.92LysGln: 3.92 ± 1.164
4.312LysArg: 4.312 ± 0.892
3.136LysSer: 3.136 ± 1.509
2.352LysThr: 2.352 ± 0.937
5.488LysVal: 5.488 ± 1.533
1.568LysTrp: 1.568 ± 0.433
1.96LysTyr: 1.96 ± 0.59
0.0LysXaa: 0.0 ± 0.0
Leu
5.096LeuAla: 5.096 ± 1.2
1.96LeuCys: 1.96 ± 0.76
9.408LeuAsp: 9.408 ± 1.74
3.136LeuGlu: 3.136 ± 1.046
5.096LeuPhe: 5.096 ± 0.888
5.88LeuGly: 5.88 ± 0.88
3.528LeuHis: 3.528 ± 1.676
5.096LeuIle: 5.096 ± 2.375
3.92LeuLys: 3.92 ± 1.787
7.84LeuLeu: 7.84 ± 1.143
0.784LeuMet: 0.784 ± 0.605
3.528LeuAsn: 3.528 ± 1.24
5.096LeuPro: 5.096 ± 1.546
5.88LeuGln: 5.88 ± 1.324
4.704LeuArg: 4.704 ± 1.584
8.624LeuSer: 8.624 ± 1.377
3.528LeuThr: 3.528 ± 1.743
5.488LeuVal: 5.488 ± 1.13
0.0LeuTrp: 0.0 ± 0.0
4.704LeuTyr: 4.704 ± 1.7
0.0LeuXaa: 0.0 ± 0.0
Met
1.568MetAla: 1.568 ± 0.933
0.784MetCys: 0.784 ± 0.737
1.176MetAsp: 1.176 ± 0.376
1.568MetGlu: 1.568 ± 0.701
0.392MetPhe: 0.392 ± 0.325
1.176MetGly: 1.176 ± 1.056
0.392MetHis: 0.392 ± 0.302
0.392MetIle: 0.392 ± 0.325
0.392MetLys: 0.392 ± 0.325
3.136MetLeu: 3.136 ± 1.34
0.0MetMet: 0.0 ± 0.0
1.568MetAsn: 1.568 ± 0.705
0.784MetPro: 0.784 ± 0.651
0.0MetGln: 0.0 ± 0.0
0.0MetArg: 0.0 ± 0.0
1.176MetSer: 1.176 ± 0.506
0.0MetThr: 0.0 ± 0.0
1.176MetVal: 1.176 ± 0.379
0.784MetTrp: 0.784 ± 0.648
0.392MetTyr: 0.392 ± 0.324
0.0MetXaa: 0.0 ± 0.0
Asn
1.96AsnAla: 1.96 ± 0.396
2.352AsnCys: 2.352 ± 1.039
1.176AsnAsp: 1.176 ± 0.661
3.136AsnGlu: 3.136 ± 1.092
1.568AsnPhe: 1.568 ± 0.705
3.92AsnGly: 3.92 ± 0.779
0.0AsnHis: 0.0 ± 0.0
2.352AsnIle: 2.352 ± 0.857
3.528AsnLys: 3.528 ± 1.034
2.352AsnLeu: 2.352 ± 0.873
2.352AsnMet: 2.352 ± 1.217
1.568AsnAsn: 1.568 ± 1.069
1.96AsnPro: 1.96 ± 0.567
2.352AsnGln: 2.352 ± 0.498
1.96AsnArg: 1.96 ± 0.774
4.312AsnSer: 4.312 ± 1.128
4.312AsnThr: 4.312 ± 1.249
2.352AsnVal: 2.352 ± 0.759
1.176AsnTrp: 1.176 ± 0.72
0.0AsnTyr: 0.0 ± 0.0
0.0AsnXaa: 0.0 ± 0.0
Pro
4.704ProAla: 4.704 ± 0.996
1.96ProCys: 1.96 ± 0.76
3.528ProAsp: 3.528 ± 1.481
3.136ProGlu: 3.136 ± 1.11
2.744ProPhe: 2.744 ± 0.98
2.744ProGly: 2.744 ± 1.114
1.96ProHis: 1.96 ± 1.953
2.352ProIle: 2.352 ± 0.66
3.528ProLys: 3.528 ± 0.904
6.272ProLeu: 6.272 ± 1.091
0.0ProMet: 0.0 ± 0.0
2.352ProAsn: 2.352 ± 0.96
10.192ProPro: 10.192 ± 5.476
1.568ProGln: 1.568 ± 0.964
2.352ProArg: 2.352 ± 1.105
7.448ProSer: 7.448 ± 2.647
2.744ProThr: 2.744 ± 1.426
3.92ProVal: 3.92 ± 1.757
0.784ProTrp: 0.784 ± 0.743
1.568ProTyr: 1.568 ± 1.197
0.0ProXaa: 0.0 ± 0.0
Gln
1.96GlnAla: 1.96 ± 0.59
0.392GlnCys: 0.392 ± 0.643
3.528GlnAsp: 3.528 ± 0.518
2.352GlnGlu: 2.352 ± 1.045
2.744GlnPhe: 2.744 ± 0.896
1.96GlnGly: 1.96 ± 0.788
0.784GlnHis: 0.784 ± 1.286
1.96GlnIle: 1.96 ± 0.728
2.744GlnLys: 2.744 ± 0.733
5.488GlnLeu: 5.488 ± 1.442
1.96GlnMet: 1.96 ± 1.081
1.96GlnAsn: 1.96 ± 0.661
3.528GlnPro: 3.528 ± 0.585
1.96GlnGln: 1.96 ± 1.29
1.96GlnArg: 1.96 ± 1.181
1.568GlnSer: 1.568 ± 0.63
2.744GlnThr: 2.744 ± 1.288
1.96GlnVal: 1.96 ± 0.597
1.568GlnTrp: 1.568 ± 0.969
0.784GlnTyr: 0.784 ± 0.444
0.0GlnXaa: 0.0 ± 0.0
Arg
5.096ArgAla: 5.096 ± 0.733
1.568ArgCys: 1.568 ± 0.998
0.392ArgAsp: 0.392 ± 0.452
2.352ArgGlu: 2.352 ± 0.574
1.176ArgPhe: 1.176 ± 0.651
3.136ArgGly: 3.136 ± 0.508
3.92ArgHis: 3.92 ± 1.181
1.176ArgIle: 1.176 ± 0.65
3.528ArgLys: 3.528 ± 0.735
6.272ArgLeu: 6.272 ± 0.951
0.784ArgMet: 0.784 ± 0.651
1.96ArgAsn: 1.96 ± 0.94
3.528ArgPro: 3.528 ± 1.603
2.352ArgGln: 2.352 ± 0.605
3.92ArgArg: 3.92 ± 1.317
1.96ArgSer: 1.96 ± 0.777
2.744ArgThr: 2.744 ± 0.638
1.96ArgVal: 1.96 ± 0.661
0.0ArgTrp: 0.0 ± 0.0
0.784ArgTyr: 0.784 ± 0.444
0.0ArgXaa: 0.0 ± 0.0
Ser
2.352SerAla: 2.352 ± 0.891
0.392SerCys: 0.392 ± 0.325
4.312SerAsp: 4.312 ± 1.617
5.88SerGlu: 5.88 ± 1.387
5.096SerPhe: 5.096 ± 1.109
5.88SerGly: 5.88 ± 1.691
2.744SerHis: 2.744 ± 1.391
3.92SerIle: 3.92 ± 1.203
3.136SerLys: 3.136 ± 1.596
8.624SerLeu: 8.624 ± 1.76
0.784SerMet: 0.784 ± 0.647
3.136SerAsn: 3.136 ± 1.273
4.312SerPro: 4.312 ± 1.284
2.352SerGln: 2.352 ± 0.867
3.528SerArg: 3.528 ± 0.943
9.016SerSer: 9.016 ± 3.61
6.664SerThr: 6.664 ± 2.29
4.312SerVal: 4.312 ± 0.969
1.176SerTrp: 1.176 ± 0.584
1.176SerTyr: 1.176 ± 0.434
0.0SerXaa: 0.0 ± 0.0
Thr
3.528ThrAla: 3.528 ± 0.943
1.96ThrCys: 1.96 ± 0.734
3.528ThrAsp: 3.528 ± 0.778
4.312ThrGlu: 4.312 ± 0.795
3.136ThrPhe: 3.136 ± 0.641
4.704ThrGly: 4.704 ± 0.658
0.784ThrHis: 0.784 ± 0.408
2.744ThrIle: 2.744 ± 0.711
3.136ThrLys: 3.136 ± 1.727
4.312ThrLeu: 4.312 ± 1.76
0.392ThrMet: 0.392 ± 0.483
1.568ThrAsn: 1.568 ± 0.666
4.312ThrPro: 4.312 ± 1.218
4.312ThrGln: 4.312 ± 1.942
2.352ThrArg: 2.352 ± 0.949
7.056ThrSer: 7.056 ± 2.144
3.92ThrThr: 3.92 ± 0.623
7.84ThrVal: 7.84 ± 2.573
1.568ThrTrp: 1.568 ± 0.889
1.568ThrTyr: 1.568 ± 0.781
0.0ThrXaa: 0.0 ± 0.0
Val
3.136ValAla: 3.136 ± 1.092
1.568ValCys: 1.568 ± 1.19
4.312ValAsp: 4.312 ± 1.12
4.312ValGlu: 4.312 ± 1.571
1.96ValPhe: 1.96 ± 0.597
4.704ValGly: 4.704 ± 0.81
0.784ValHis: 0.784 ± 0.357
1.96ValIle: 1.96 ± 0.577
2.352ValLys: 2.352 ± 0.713
5.488ValLeu: 5.488 ± 0.937
0.0ValMet: 0.0 ± 0.0
2.352ValAsn: 2.352 ± 0.884
4.312ValPro: 4.312 ± 1.037
1.96ValGln: 1.96 ± 1.052
3.528ValArg: 3.528 ± 1.034
6.664ValSer: 6.664 ± 1.913
5.488ValThr: 5.488 ± 1.655
4.312ValVal: 4.312 ± 0.87
0.392ValTrp: 0.392 ± 0.324
0.784ValTyr: 0.784 ± 0.467
0.0ValXaa: 0.0 ± 0.0
Trp
1.176TrpAla: 1.176 ± 0.661
0.0TrpCys: 0.0 ± 0.0
0.392TrpAsp: 0.392 ± 0.325
0.784TrpGlu: 0.784 ± 0.737
0.784TrpPhe: 0.784 ± 0.444
0.784TrpGly: 0.784 ± 0.465
0.0TrpHis: 0.0 ± 0.0
1.96TrpIle: 1.96 ± 0.935
1.96TrpLys: 1.96 ± 0.569
1.96TrpLeu: 1.96 ± 0.956
0.0TrpMet: 0.0 ± 0.0
0.784TrpAsn: 0.784 ± 0.408
0.0TrpPro: 0.0 ± 0.0
0.392TrpGln: 0.392 ± 0.452
0.784TrpArg: 0.784 ± 0.752
0.784TrpSer: 0.784 ± 0.634
1.176TrpThr: 1.176 ± 0.937
1.176TrpVal: 1.176 ± 0.783
0.0TrpTrp: 0.0 ± 0.0
0.392TrpTyr: 0.392 ± 0.325
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.352TyrAla: 2.352 ± 0.881
0.392TyrCys: 0.392 ± 0.325
1.568TyrAsp: 1.568 ± 0.701
1.568TyrGlu: 1.568 ± 0.607
0.784TyrPhe: 0.784 ± 0.444
1.176TyrGly: 1.176 ± 0.617
0.784TyrHis: 0.784 ± 0.39
0.392TyrIle: 0.392 ± 0.533
0.784TyrLys: 0.784 ± 0.42
1.568TyrLeu: 1.568 ± 0.609
0.392TyrMet: 0.392 ± 0.276
1.176TyrAsn: 1.176 ± 0.651
1.96TyrPro: 1.96 ± 0.848
0.392TyrGln: 0.392 ± 0.324
1.96TyrArg: 1.96 ± 0.996
2.744TyrSer: 2.744 ± 1.185
1.96TyrThr: 1.96 ± 0.45
1.568TyrVal: 1.568 ± 0.709
1.176TyrTrp: 1.176 ± 0.972
1.568TyrTyr: 1.568 ± 0.996
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (2552 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski