Amino acid dipepetide frequency for Oryza glumipatula

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	14.885AlaAla: 14.885 ± 0.06	1.576AlaCys: 1.576 ± 0.011	4.218AlaAsp: 4.218 ± 0.017	4.844AlaGlu: 4.844 ± 0.022	2.905AlaPhe: 2.905 ± 0.015	6.726AlaGly: 6.726 ± 0.029	1.748AlaHis: 1.748 ± 0.012	3.714AlaIle: 3.714 ± 0.017	3.618AlaLys: 3.618 ± 0.015	7.679AlaLeu: 7.679 ± 0.027	2.326AlaMet: 2.326 ± 0.011	2.547AlaAsn: 2.547 ± 0.012	4.728AlaPro: 4.728 ± 0.024	2.382AlaGln: 2.382 ± 0.013	5.476AlaArg: 5.476 ± 0.022	7.458AlaSer: 7.458 ± 0.026	4.868AlaThr: 4.868 ± 0.018	6.784AlaVal: 6.784 ± 0.025	1.018AlaTrp: 1.018 ± 0.01	1.892AlaTyr: 1.892 ± 0.011	0.002AlaXaa: 0.002 ± 0.0
Cys	1.362CysAla: 1.362 ± 0.01	0.636CysCys: 0.636 ± 0.007	0.909CysAsp: 0.909 ± 0.007	0.823CysGlu: 0.823 ± 0.007	0.827CysPhe: 0.827 ± 0.007	1.574CysGly: 1.574 ± 0.013	0.55CysHis: 0.55 ± 0.005	0.935CysIle: 0.935 ± 0.008	0.956CysLys: 0.956 ± 0.009	1.85CysLeu: 1.85 ± 0.011	0.461CysMet: 0.461 ± 0.005	0.754CysAsn: 0.754 ± 0.008	1.039CysPro: 1.039 ± 0.009	0.586CysGln: 0.586 ± 0.006	1.365CysArg: 1.365 ± 0.01	1.936CysSer: 1.936 ± 0.013	0.945CysThr: 0.945 ± 0.008	1.134CysVal: 1.134 ± 0.01	0.269CysTrp: 0.269 ± 0.004	0.537CysTyr: 0.537 ± 0.006	0.0CysXaa: 0.0 ± 0.0
Asp	4.734AspAla: 4.734 ± 0.02	0.928AspCys: 0.928 ± 0.008	4.444AspAsp: 4.444 ± 0.028	3.912AspGlu: 3.912 ± 0.022	1.943AspPhe: 1.943 ± 0.012	4.797AspGly: 4.797 ± 0.021	1.303AspHis: 1.303 ± 0.01	2.699AspIle: 2.699 ± 0.014	2.368AspLys: 2.368 ± 0.013	4.868AspLeu: 4.868 ± 0.02	1.415AspMet: 1.415 ± 0.009	1.823AspAsn: 1.823 ± 0.013	2.602AspPro: 2.602 ± 0.013	1.593AspGln: 1.593 ± 0.011	2.689AspArg: 2.689 ± 0.015	3.667AspSer: 3.667 ± 0.018	2.212AspThr: 2.212 ± 0.013	3.754AspVal: 3.754 ± 0.017	0.675AspTrp: 0.675 ± 0.006	1.366AspTyr: 1.366 ± 0.01	0.001AspXaa: 0.001 ± 0.0
Glu	5.302GluAla: 5.302 ± 0.02	0.868GluCys: 0.868 ± 0.01	3.628GluAsp: 3.628 ± 0.022	5.913GluGlu: 5.913 ± 0.032	1.98GluPhe: 1.98 ± 0.011	3.715GluGly: 3.715 ± 0.017	1.404GluHis: 1.404 ± 0.011	2.969GluIle: 2.969 ± 0.015	3.671GluLys: 3.671 ± 0.022	5.9GluLeu: 5.9 ± 0.023	1.691GluMet: 1.691 ± 0.01	2.226GluAsn: 2.226 ± 0.014	2.235GluPro: 2.235 ± 0.013	2.186GluGln: 2.186 ± 0.012	3.652GluArg: 3.652 ± 0.018	3.715GluSer: 3.715 ± 0.019	2.538GluThr: 2.538 ± 0.016	4.03GluVal: 4.03 ± 0.019	0.756GluTrp: 0.756 ± 0.007	1.453GluTyr: 1.453 ± 0.01	0.001GluXaa: 0.001 ± 0.0
Phe	2.739PheAla: 2.739 ± 0.014	0.786PheCys: 0.786 ± 0.007	2.104PheAsp: 2.104 ± 0.013	1.778PheGlu: 1.778 ± 0.01	1.675PhePhe: 1.675 ± 0.013	2.806PheGly: 2.806 ± 0.016	0.992PheHis: 0.992 ± 0.007	1.608PheIle: 1.608 ± 0.013	1.456PheLys: 1.456 ± 0.01	3.842PheLeu: 3.842 ± 0.018	0.851PheMet: 0.851 ± 0.008	1.291PheAsn: 1.291 ± 0.009	1.889PhePro: 1.889 ± 0.011	1.228PheGln: 1.228 ± 0.009	2.016PheArg: 2.016 ± 0.012	3.299PheSer: 3.299 ± 0.017	1.733PheThr: 1.733 ± 0.011	2.575PheVal: 2.575 ± 0.011	0.501PheTrp: 0.501 ± 0.005	1.026PheTyr: 1.026 ± 0.008	0.001PheXaa: 0.001 ± 0.0
Gly	6.125GlyAla: 6.125 ± 0.026	1.537GlyCys: 1.537 ± 0.01	4.271GlyAsp: 4.271 ± 0.02	4.282GlyGlu: 4.282 ± 0.019	2.787GlyPhe: 2.787 ± 0.017	9.433GlyGly: 9.433 ± 0.073	1.794GlyHis: 1.794 ± 0.011	3.145GlyIle: 3.145 ± 0.016	3.548GlyLys: 3.548 ± 0.017	5.803GlyLeu: 5.803 ± 0.022	1.672GlyMet: 1.672 ± 0.012	2.733GlyAsn: 2.733 ± 0.015	2.624GlyPro: 2.624 ± 0.013	2.118GlyGln: 2.118 ± 0.015	5.003GlyArg: 5.003 ± 0.024	6.097GlySer: 6.097 ± 0.029	3.317GlyThr: 3.317 ± 0.014	4.78GlyVal: 4.78 ± 0.018	1.034GlyTrp: 1.034 ± 0.009	2.025GlyTyr: 2.025 ± 0.015	0.002GlyXaa: 0.002 ± 0.0
His	1.956HisAla: 1.956 ± 0.011	0.568HisCys: 0.568 ± 0.006	1.29HisAsp: 1.29 ± 0.009	1.235HisGlu: 1.235 ± 0.01	0.912HisPhe: 0.912 ± 0.008	2.179HisGly: 2.179 ± 0.012	1.186HisHis: 1.186 ± 0.013	1.101HisIle: 1.101 ± 0.007	0.987HisLys: 0.987 ± 0.008	2.647HisLeu: 2.647 ± 0.014	0.567HisMet: 0.567 ± 0.006	0.824HisAsn: 0.824 ± 0.007	1.601HisPro: 1.601 ± 0.011	0.962HisGln: 0.962 ± 0.008	1.86HisArg: 1.86 ± 0.012	1.725HisSer: 1.725 ± 0.011	0.994HisThr: 0.994 ± 0.007	1.628HisVal: 1.628 ± 0.01	0.303HisTrp: 0.303 ± 0.004	0.646HisTyr: 0.646 ± 0.006	0.0HisXaa: 0.0 ± 0.0
Ile	3.55IleAla: 3.55 ± 0.016	0.997IleCys: 0.997 ± 0.009	2.546IleAsp: 2.546 ± 0.014	2.487IleGlu: 2.487 ± 0.014	1.726IlePhe: 1.726 ± 0.011	3.064IleGly: 3.064 ± 0.015	1.163IleHis: 1.163 ± 0.009	2.272IleIle: 2.272 ± 0.013	2.248IleLys: 2.248 ± 0.011	4.381IleLeu: 4.381 ± 0.019	1.004IleMet: 1.004 ± 0.008	1.706IleAsn: 1.706 ± 0.011	2.527IlePro: 2.527 ± 0.018	1.567IleGln: 1.567 ± 0.012	2.433IleArg: 2.433 ± 0.014	3.998IleSer: 3.998 ± 0.019	2.282IleThr: 2.282 ± 0.013	3.032IleVal: 3.032 ± 0.013	0.618IleTrp: 0.618 ± 0.006	1.247IleTyr: 1.247 ± 0.01	0.001IleXaa: 0.001 ± 0.0
Lys	3.664LysAla: 3.664 ± 0.015	0.825LysCys: 0.825 ± 0.008	2.618LysAsp: 2.618 ± 0.018	3.588LysGlu: 3.588 ± 0.02	1.58LysPhe: 1.58 ± 0.011	2.988LysGly: 2.988 ± 0.016	1.152LysHis: 1.152 ± 0.008	2.542LysIle: 2.542 ± 0.015	3.56LysLys: 3.56 ± 0.023	4.885LysLeu: 4.885 ± 0.022	1.253LysMet: 1.253 ± 0.009	1.984LysAsn: 1.984 ± 0.012	2.278LysPro: 2.278 ± 0.014	1.847LysGln: 1.847 ± 0.014	3.084LysArg: 3.084 ± 0.014	3.393LysSer: 3.393 ± 0.018	2.199LysThr: 2.199 ± 0.012	3.123LysVal: 3.123 ± 0.015	0.595LysTrp: 0.595 ± 0.007	1.293LysTyr: 1.293 ± 0.01	0.001LysXaa: 0.001 ± 0.0
Leu	8.127LeuAla: 8.127 ± 0.031	1.85LeuCys: 1.85 ± 0.012	5.076LeuAsp: 5.076 ± 0.021	5.669LeuGlu: 5.669 ± 0.025	3.373LeuPhe: 3.373 ± 0.016	5.837LeuGly: 5.837 ± 0.021	2.82LeuHis: 2.82 ± 0.013	3.73LeuIle: 3.73 ± 0.018	4.674LeuLys: 4.674 ± 0.024	10.681LeuLeu: 10.681 ± 0.036	2.001LeuMet: 2.001 ± 0.012	3.071LeuAsn: 3.071 ± 0.017	6.029LeuPro: 6.029 ± 0.02	4.037LeuGln: 4.037 ± 0.02	6.477LeuArg: 6.477 ± 0.024	7.884LeuSer: 7.884 ± 0.037	4.281LeuThr: 4.281 ± 0.022	6.449LeuVal: 6.449 ± 0.022	1.14LeuTrp: 1.14 ± 0.01	2.268LeuTyr: 2.268 ± 0.013	0.001LeuXaa: 0.001 ± 0.0
Met	2.827MetAla: 2.827 ± 0.013	0.346MetCys: 0.346 ± 0.005	1.473MetAsp: 1.473 ± 0.01	1.87MetGlu: 1.87 ± 0.011	0.767MetPhe: 0.767 ± 0.007	1.52MetGly: 1.52 ± 0.01	0.579MetHis: 0.579 ± 0.006	1.003MetIle: 1.003 ± 0.009	1.28MetLys: 1.28 ± 0.009	2.231MetLeu: 2.231 ± 0.013	0.68MetMet: 0.68 ± 0.007	0.854MetAsn: 0.854 ± 0.008	1.275MetPro: 1.275 ± 0.009	0.896MetGln: 0.896 ± 0.007	1.388MetArg: 1.388 ± 0.011	1.777MetSer: 1.777 ± 0.009	1.044MetThr: 1.044 ± 0.008	1.717MetVal: 1.717 ± 0.009	0.3MetTrp: 0.3 ± 0.004	0.578MetTyr: 0.578 ± 0.006	0.0MetXaa: 0.0 ± 0.0
Asn	2.585AsnAla: 2.585 ± 0.014	0.727AsnCys: 0.727 ± 0.007	1.707AsnAsp: 1.707 ± 0.011	1.83AsnGlu: 1.83 ± 0.011	1.426AsnPhe: 1.426 ± 0.011	2.838AsnGly: 2.838 ± 0.014	0.913AsnHis: 0.913 ± 0.008	2.009AsnIle: 2.009 ± 0.012	1.804AsnLys: 1.804 ± 0.014	3.807AsnLeu: 3.807 ± 0.027	0.989AsnMet: 0.989 ± 0.008	1.731AsnAsn: 1.731 ± 0.016	1.859AsnPro: 1.859 ± 0.011	1.258AsnGln: 1.258 ± 0.01	1.757AsnArg: 1.757 ± 0.009	2.915AsnSer: 2.915 ± 0.016	1.686AsnThr: 1.686 ± 0.01	2.231AsnVal: 2.231 ± 0.013	0.441AsnTrp: 0.441 ± 0.005	0.979AsnTyr: 0.979 ± 0.008	0.0AsnXaa: 0.0 ± 0.0
Pro	5.283ProAla: 5.283 ± 0.023	0.99ProCys: 0.99 ± 0.007	2.716ProAsp: 2.716 ± 0.014	3.18ProGlu: 3.18 ± 0.016	1.855ProPhe: 1.855 ± 0.01	3.202ProGly: 3.202 ± 0.016	1.304ProHis: 1.304 ± 0.01	1.951ProIle: 1.951 ± 0.011	2.233ProLys: 2.233 ± 0.014	4.704ProLeu: 4.704 ± 0.019	1.09ProMet: 1.09 ± 0.008	1.852ProAsn: 1.852 ± 0.01	6.203ProPro: 6.203 ± 0.043	1.743ProGln: 1.743 ± 0.013	3.532ProArg: 3.532 ± 0.017	5.52ProSer: 5.52 ± 0.026	2.829ProThr: 2.829 ± 0.013	3.398ProVal: 3.398 ± 0.015	0.704ProTrp: 0.704 ± 0.007	1.261ProTyr: 1.261 ± 0.009	0.001ProXaa: 0.001 ± 0.0
Gln	2.561GlnAla: 2.561 ± 0.015	0.598GlnCys: 0.598 ± 0.006	1.573GlnAsp: 1.573 ± 0.012	2.216GlnGlu: 2.216 ± 0.014	1.145GlnPhe: 1.145 ± 0.009	2.112GlnGly: 2.112 ± 0.011	0.989GlnHis: 0.989 ± 0.008	1.606GlnIle: 1.606 ± 0.009	1.814GlnLys: 1.814 ± 0.013	3.508GlnLeu: 3.508 ± 0.02	0.917GlnMet: 0.917 ± 0.008	1.318GlnAsn: 1.318 ± 0.01	1.91GlnPro: 1.91 ± 0.015	2.158GlnGln: 2.158 ± 0.027	2.259GlnArg: 2.259 ± 0.014	2.398GlnSer: 2.398 ± 0.014	1.441GlnThr: 1.441 ± 0.01	2.117GlnVal: 2.117 ± 0.011	0.461GlnTrp: 0.461 ± 0.005	0.867GlnTyr: 0.867 ± 0.008	0.0GlnXaa: 0.0 ± 0.0
Arg	5.083ArgAla: 5.083 ± 0.023	1.407ArgCys: 1.407 ± 0.009	2.972ArgAsp: 2.972 ± 0.017	3.538ArgGlu: 3.538 ± 0.019	2.211ArgPhe: 2.211 ± 0.014	4.478ArgGly: 4.478 ± 0.021	1.767ArgHis: 1.767 ± 0.012	2.674ArgIle: 2.674 ± 0.014	3.289ArgLys: 3.289 ± 0.016	6.121ArgLeu: 6.121 ± 0.026	1.557ArgMet: 1.557 ± 0.01	2.124ArgAsn: 2.124 ± 0.012	3.392ArgPro: 3.392 ± 0.017	2.093ArgGln: 2.093 ± 0.012	7.453ArgArg: 7.453 ± 0.068	5.028ArgSer: 5.028 ± 0.021	2.68ArgThr: 2.68 ± 0.014	3.804ArgVal: 3.804 ± 0.017	1.121ArgTrp: 1.121 ± 0.008	1.578ArgTyr: 1.578 ± 0.011	0.002ArgXaa: 0.002 ± 0.0
Ser	6.498SerAla: 6.498 ± 0.022	1.783SerCys: 1.783 ± 0.012	4.018SerAsp: 4.018 ± 0.017	3.986SerGlu: 3.986 ± 0.02	3.293SerPhe: 3.293 ± 0.016	6.082SerGly: 6.082 ± 0.033	1.845SerHis: 1.845 ± 0.01	3.743SerIle: 3.743 ± 0.018	3.744SerLys: 3.744 ± 0.02	7.916SerLeu: 7.916 ± 0.027	2.031SerMet: 2.031 ± 0.012	3.144SerAsn: 3.144 ± 0.017	5.179SerPro: 5.179 ± 0.022	2.478SerGln: 2.478 ± 0.015	4.857SerArg: 4.857 ± 0.022	10.638SerSer: 10.638 ± 0.043	4.542SerThr: 4.542 ± 0.022	4.818SerVal: 4.818 ± 0.019	1.162SerTrp: 1.162 ± 0.009	2.1SerTyr: 2.1 ± 0.011	0.001SerXaa: 0.001 ± 0.0
Thr	4.479ThrAla: 4.479 ± 0.018	0.954ThrCys: 0.954 ± 0.009	2.268ThrAsp: 2.268 ± 0.015	2.534ThrGlu: 2.534 ± 0.012	1.732ThrPhe: 1.732 ± 0.011	3.494ThrGly: 3.494 ± 0.016	0.98ThrHis: 0.98 ± 0.007	2.345ThrIle: 2.345 ± 0.013	2.208ThrLys: 2.208 ± 0.011	4.21ThrLeu: 4.21 ± 0.019	1.272ThrMet: 1.272 ± 0.008	1.782ThrAsn: 1.782 ± 0.01	2.818ThrPro: 2.818 ± 0.016	1.332ThrGln: 1.332 ± 0.01	2.661ThrArg: 2.661 ± 0.013	4.341ThrSer: 4.341 ± 0.018	3.263ThrThr: 3.263 ± 0.018	3.47ThrVal: 3.47 ± 0.015	0.62ThrTrp: 0.62 ± 0.006	1.259ThrTyr: 1.259 ± 0.01	0.001ThrXaa: 0.001 ± 0.0
Val	6.81ValAla: 6.81 ± 0.028	1.181ValCys: 1.181 ± 0.009	3.866ValAsp: 3.866 ± 0.014	4.102ValGlu: 4.102 ± 0.018	2.485ValPhe: 2.485 ± 0.013	4.536ValGly: 4.536 ± 0.017	1.667ValHis: 1.667 ± 0.009	2.891ValIle: 2.891 ± 0.015	3.071ValLys: 3.071 ± 0.013	6.573ValLeu: 6.573 ± 0.023	1.546ValMet: 1.546 ± 0.01	2.046ValAsn: 2.046 ± 0.013	3.561ValPro: 3.561 ± 0.017	2.227ValGln: 2.227 ± 0.013	3.774ValArg: 3.774 ± 0.019	5.047ValSer: 5.047 ± 0.017	3.294ValThr: 3.294 ± 0.015	5.725ValVal: 5.725 ± 0.022	0.816ValTrp: 0.816 ± 0.008	1.756ValTyr: 1.756 ± 0.012	0.001ValXaa: 0.001 ± 0.0
Trp	0.96TrpAla: 0.96 ± 0.009	0.286TrpCys: 0.286 ± 0.004	0.654TrpAsp: 0.654 ± 0.006	0.736TrpGlu: 0.736 ± 0.007	0.486TrpPhe: 0.486 ± 0.005	0.781TrpGly: 0.781 ± 0.008	0.319TrpHis: 0.319 ± 0.004	0.628TrpIle: 0.628 ± 0.007	0.731TrpLys: 0.731 ± 0.006	1.282TrpLeu: 1.282 ± 0.009	0.38TrpMet: 0.38 ± 0.005	0.568TrpAsn: 0.568 ± 0.006	0.625TrpPro: 0.625 ± 0.006	0.464TrpGln: 0.464 ± 0.005	1.171TrpArg: 1.171 ± 0.01	1.048TrpSer: 1.048 ± 0.008	0.666TrpThr: 0.666 ± 0.006	0.799TrpVal: 0.799 ± 0.008	0.29TrpTrp: 0.29 ± 0.004	0.331TrpTyr: 0.331 ± 0.005	0.0TrpXaa: 0.0 ± 0.0
Tyr	1.861TyrAla: 1.861 ± 0.009	0.603TyrCys: 0.603 ± 0.006	1.443TyrAsp: 1.443 ± 0.01	1.31TyrGlu: 1.31 ± 0.009	1.085TyrPhe: 1.085 ± 0.008	1.965TyrGly: 1.965 ± 0.011	0.698TyrHis: 0.698 ± 0.007	1.283TyrIle: 1.283 ± 0.009	1.192TyrLys: 1.192 ± 0.01	2.587TyrLeu: 2.587 ± 0.014	0.701TyrMet: 0.701 ± 0.006	1.113TyrAsn: 1.113 ± 0.012	1.175TyrPro: 1.175 ± 0.01	0.817TyrGln: 0.817 ± 0.008	1.466TyrArg: 1.466 ± 0.01	1.973TyrSer: 1.973 ± 0.011	1.206TyrThr: 1.206 ± 0.009	1.606TyrVal: 1.606 ± 0.011	0.38TyrTrp: 0.38 ± 0.005	0.909TyrTyr: 0.909 ± 0.009	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.002XaaAla: 0.002 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.001XaaAsp: 0.001 ± 0.0	0.001XaaGlu: 0.001 ± 0.0	0.001XaaPhe: 0.001 ± 0.0	0.002XaaGly: 0.002 ± 0.0	0.001XaaHis: 0.001 ± 0.0	0.001XaaIle: 0.001 ± 0.0	0.0XaaLys: 0.0 ± 0.0	0.002XaaLeu: 0.002 ± 0.0	0.001XaaMet: 0.001 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.001XaaPro: 0.001 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.002XaaArg: 0.002 ± 0.0	0.001XaaSer: 0.001 ± 0.0	0.001XaaThr: 0.001 ± 0.0	0.001XaaVal: 0.001 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.012XaaXaa: 0.012 ± 0.003

Statistics based on 44731 proteins (18292435 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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