Amino acid dipepetide frequency for Beet soil-borne virus

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
5.744AlaAla: 5.744 ± 1.113
1.026AlaCys: 1.026 ± 0.336
3.487AlaAsp: 3.487 ± 0.565
4.308AlaGlu: 4.308 ± 0.932
2.256AlaPhe: 2.256 ± 1.083
2.051AlaGly: 2.051 ± 1.282
1.846AlaHis: 1.846 ± 0.392
3.897AlaIle: 3.897 ± 1.12
5.744AlaLys: 5.744 ± 0.463
8.0AlaLeu: 8.0 ± 1.314
1.846AlaMet: 1.846 ± 0.443
2.462AlaAsn: 2.462 ± 0.272
1.231AlaPro: 1.231 ± 0.612
1.231AlaGln: 1.231 ± 0.383
3.897AlaArg: 3.897 ± 0.628
3.692AlaSer: 3.692 ± 0.762
4.103AlaThr: 4.103 ± 1.003
4.718AlaVal: 4.718 ± 0.878
1.026AlaTrp: 1.026 ± 0.274
1.846AlaTyr: 1.846 ± 0.733
0.0AlaXaa: 0.0 ± 0.0
Cys
1.641CysAla: 1.641 ± 0.589
1.231CysCys: 1.231 ± 0.607
1.641CysAsp: 1.641 ± 0.517
1.026CysGlu: 1.026 ± 0.338
1.436CysPhe: 1.436 ± 0.316
2.051CysGly: 2.051 ± 0.327
0.0CysHis: 0.0 ± 0.0
0.615CysIle: 0.615 ± 0.176
1.026CysLys: 1.026 ± 0.429
0.41CysLeu: 0.41 ± 0.364
0.0CysMet: 0.0 ± 0.0
0.821CysAsn: 0.821 ± 0.427
0.615CysPro: 0.615 ± 0.304
1.436CysGln: 1.436 ± 0.469
0.821CysArg: 0.821 ± 0.482
0.821CysSer: 0.821 ± 0.772
0.821CysThr: 0.821 ± 0.529
1.846CysVal: 1.846 ± 0.355
0.0CysTrp: 0.0 ± 0.0
0.615CysTyr: 0.615 ± 0.277
0.0CysXaa: 0.0 ± 0.0
Asp
3.077AspAla: 3.077 ± 0.561
0.821AspCys: 0.821 ± 0.339
5.744AspAsp: 5.744 ± 1.268
4.718AspGlu: 4.718 ± 0.441
3.282AspPhe: 3.282 ± 1.068
3.487AspGly: 3.487 ± 0.631
0.41AspHis: 0.41 ± 0.179
5.744AspIle: 5.744 ± 1.47
4.513AspLys: 4.513 ± 0.538
6.974AspLeu: 6.974 ± 0.841
1.641AspMet: 1.641 ± 0.478
4.308AspAsn: 4.308 ± 0.585
3.077AspPro: 3.077 ± 0.861
1.641AspGln: 1.641 ± 0.761
2.872AspArg: 2.872 ± 1.285
5.538AspSer: 5.538 ± 0.631
3.077AspThr: 3.077 ± 1.271
6.359AspVal: 6.359 ± 0.496
0.615AspTrp: 0.615 ± 0.265
2.667AspTyr: 2.667 ± 0.398
0.0AspXaa: 0.0 ± 0.0
Glu
6.359GluAla: 6.359 ± 1.102
1.231GluCys: 1.231 ± 0.425
7.385GluAsp: 7.385 ± 1.209
5.744GluGlu: 5.744 ± 1.282
2.462GluPhe: 2.462 ± 0.639
5.333GluGly: 5.333 ± 0.799
1.641GluHis: 1.641 ± 0.458
4.923GluIle: 4.923 ± 0.579
5.744GluLys: 5.744 ± 0.516
4.308GluLeu: 4.308 ± 0.466
3.077GluMet: 3.077 ± 0.689
2.051GluAsn: 2.051 ± 0.493
0.41GluPro: 0.41 ± 0.179
1.436GluGln: 1.436 ± 0.348
4.923GluArg: 4.923 ± 0.38
6.154GluSer: 6.154 ± 0.667
3.487GluThr: 3.487 ± 0.87
4.513GluVal: 4.513 ± 1.586
0.205GluTrp: 0.205 ± 0.32
2.667GluTyr: 2.667 ± 0.832
0.205GluXaa: 0.205 ± 0.225
Phe
2.256PheAla: 2.256 ± 0.693
1.231PheCys: 1.231 ± 0.383
3.692PheAsp: 3.692 ± 0.822
1.846PheGlu: 1.846 ± 0.724
1.846PhePhe: 1.846 ± 0.48
3.282PheGly: 3.282 ± 0.778
1.641PheHis: 1.641 ± 0.714
2.256PheIle: 2.256 ± 0.358
1.641PheLys: 1.641 ± 0.302
5.538PheLeu: 5.538 ± 0.7
1.436PheMet: 1.436 ± 0.44
1.026PheAsn: 1.026 ± 0.418
1.846PhePro: 1.846 ± 0.56
1.231PheGln: 1.231 ± 0.43
0.821PheArg: 0.821 ± 0.199
4.103PheSer: 4.103 ± 1.036
3.897PheThr: 3.897 ± 0.748
2.667PheVal: 2.667 ± 0.295
0.205PheTrp: 0.205 ± 0.318
3.077PheTyr: 3.077 ± 0.584
0.0PheXaa: 0.0 ± 0.0
Gly
3.077GlyAla: 3.077 ± 0.594
0.821GlyCys: 0.821 ± 0.572
4.718GlyAsp: 4.718 ± 0.557
4.308GlyGlu: 4.308 ± 0.927
2.051GlyPhe: 2.051 ± 0.508
3.692GlyGly: 3.692 ± 1.355
0.615GlyHis: 0.615 ± 0.304
2.051GlyIle: 2.051 ± 0.209
6.154GlyLys: 6.154 ± 1.593
3.692GlyLeu: 3.692 ± 1.063
1.436GlyMet: 1.436 ± 0.406
2.872GlyAsn: 2.872 ± 0.806
0.821GlyPro: 0.821 ± 0.199
1.026GlyGln: 1.026 ± 0.274
5.333GlyArg: 5.333 ± 0.908
4.103GlySer: 4.103 ± 0.774
2.256GlyThr: 2.256 ± 0.884
3.282GlyVal: 3.282 ± 0.653
0.41GlyTrp: 0.41 ± 0.329
1.641GlyTyr: 1.641 ± 0.57
0.0GlyXaa: 0.0 ± 0.0
His
2.051HisAla: 2.051 ± 0.571
1.436HisCys: 1.436 ± 0.602
0.41HisAsp: 0.41 ± 0.201
1.846HisGlu: 1.846 ± 0.276
1.641HisPhe: 1.641 ± 0.714
1.026HisGly: 1.026 ± 0.453
0.205HisHis: 0.205 ± 0.318
1.231HisIle: 1.231 ± 0.53
2.256HisLys: 2.256 ± 0.449
0.615HisLeu: 0.615 ± 0.277
0.0HisMet: 0.0 ± 0.0
0.41HisAsn: 0.41 ± 0.179
0.205HisPro: 0.205 ± 0.318
0.821HisGln: 0.821 ± 0.229
1.026HisArg: 1.026 ± 0.338
1.846HisSer: 1.846 ± 0.396
0.205HisThr: 0.205 ± 0.318
1.026HisVal: 1.026 ± 0.429
0.205HisTrp: 0.205 ± 0.318
0.615HisTyr: 0.615 ± 0.67
0.0HisXaa: 0.0 ± 0.0
Ile
2.667IleAla: 2.667 ± 0.574
0.821IleCys: 0.821 ± 0.235
3.077IleAsp: 3.077 ± 0.374
5.744IleGlu: 5.744 ± 0.603
1.231IlePhe: 1.231 ± 0.471
2.872IleGly: 2.872 ± 0.647
1.231IleHis: 1.231 ± 0.502
6.154IleIle: 6.154 ± 1.036
3.077IleLys: 3.077 ± 0.464
6.154IleLeu: 6.154 ± 0.626
1.641IleMet: 1.641 ± 0.465
2.462IleAsn: 2.462 ± 0.255
3.077IlePro: 3.077 ± 0.434
1.231IleGln: 1.231 ± 0.203
3.692IleArg: 3.692 ± 0.784
2.872IleSer: 2.872 ± 0.822
2.667IleThr: 2.667 ± 0.877
3.077IleVal: 3.077 ± 0.755
0.41IleTrp: 0.41 ± 0.179
1.436IleTyr: 1.436 ± 0.465
0.0IleXaa: 0.0 ± 0.0
Lys
3.487LysAla: 3.487 ± 0.533
1.231LysCys: 1.231 ± 0.279
4.103LysAsp: 4.103 ± 0.651
5.128LysGlu: 5.128 ± 1.357
4.718LysPhe: 4.718 ± 0.948
3.077LysGly: 3.077 ± 0.298
1.436LysHis: 1.436 ± 0.556
3.282LysIle: 3.282 ± 0.468
4.513LysLys: 4.513 ± 0.672
5.949LysLeu: 5.949 ± 0.632
0.41LysMet: 0.41 ± 0.179
4.513LysAsn: 4.513 ± 0.949
3.077LysPro: 3.077 ± 0.473
1.846LysGln: 1.846 ± 0.376
5.538LysArg: 5.538 ± 0.684
6.564LysSer: 6.564 ± 0.793
5.333LysThr: 5.333 ± 1.138
4.308LysVal: 4.308 ± 0.744
0.615LysTrp: 0.615 ± 0.265
1.641LysTyr: 1.641 ± 0.69
0.0LysXaa: 0.0 ± 0.0
Leu
5.744LeuAla: 5.744 ± 1.349
1.846LeuCys: 1.846 ± 0.651
3.692LeuAsp: 3.692 ± 0.879
6.359LeuGlu: 6.359 ± 0.988
4.513LeuPhe: 4.513 ± 0.84
4.308LeuGly: 4.308 ± 0.535
1.846LeuHis: 1.846 ± 0.56
2.872LeuIle: 2.872 ± 0.679
4.513LeuLys: 4.513 ± 0.961
5.949LeuLeu: 5.949 ± 2.424
2.872LeuMet: 2.872 ± 0.399
5.949LeuAsn: 5.949 ± 0.597
2.872LeuPro: 2.872 ± 0.331
4.308LeuGln: 4.308 ± 0.854
5.128LeuArg: 5.128 ± 1.484
7.385LeuSer: 7.385 ± 0.639
6.769LeuThr: 6.769 ± 2.273
3.692LeuVal: 3.692 ± 0.618
0.821LeuTrp: 0.821 ± 0.385
1.641LeuTyr: 1.641 ± 0.691
0.0LeuXaa: 0.0 ± 0.0
Met
2.667MetAla: 2.667 ± 0.733
0.41MetCys: 0.41 ± 0.368
0.615MetAsp: 0.615 ± 0.506
1.846MetGlu: 1.846 ± 0.261
1.231MetPhe: 1.231 ± 0.531
0.41MetGly: 0.41 ± 0.179
0.0MetHis: 0.0 ± 0.0
1.436MetIle: 1.436 ± 0.602
1.436MetLys: 1.436 ± 0.745
2.256MetLeu: 2.256 ± 0.3
0.615MetMet: 0.615 ± 0.176
1.641MetAsn: 1.641 ± 0.691
0.615MetPro: 0.615 ± 0.403
0.41MetGln: 0.41 ± 0.179
1.641MetArg: 1.641 ± 0.725
0.615MetSer: 0.615 ± 0.176
2.256MetThr: 2.256 ± 0.576
2.667MetVal: 2.667 ± 0.597
0.205MetTrp: 0.205 ± 0.141
1.641MetTyr: 1.641 ± 0.589
0.0MetXaa: 0.0 ± 0.0
Asn
2.256AsnAla: 2.256 ± 0.421
1.641AsnCys: 1.641 ± 0.374
3.077AsnAsp: 3.077 ± 0.752
3.077AsnGlu: 3.077 ± 0.441
3.077AsnPhe: 3.077 ± 0.924
3.897AsnGly: 3.897 ± 0.572
0.615AsnHis: 0.615 ± 0.277
2.872AsnIle: 2.872 ± 0.488
3.077AsnLys: 3.077 ± 0.928
2.667AsnLeu: 2.667 ± 0.639
1.231AsnMet: 1.231 ± 0.483
1.436AsnAsn: 1.436 ± 0.659
0.41AsnPro: 0.41 ± 0.4
1.436AsnGln: 1.436 ± 0.38
3.077AsnArg: 3.077 ± 0.49
4.718AsnSer: 4.718 ± 0.925
3.692AsnThr: 3.692 ± 0.791
3.692AsnVal: 3.692 ± 0.666
0.821AsnTrp: 0.821 ± 0.229
1.436AsnTyr: 1.436 ± 0.411
0.0AsnXaa: 0.0 ± 0.0
Pro
1.026ProAla: 1.026 ± 0.545
0.41ProCys: 0.41 ± 0.201
2.462ProAsp: 2.462 ± 0.583
2.051ProGlu: 2.051 ± 1.16
0.821ProPhe: 0.821 ± 0.339
2.872ProGly: 2.872 ± 0.716
0.615ProHis: 0.615 ± 0.265
1.231ProIle: 1.231 ± 0.545
2.256ProLys: 2.256 ± 0.802
1.231ProLeu: 1.231 ± 0.383
0.821ProMet: 0.821 ± 0.199
1.436ProAsn: 1.436 ± 0.347
1.641ProPro: 1.641 ± 0.381
1.026ProGln: 1.026 ± 0.706
2.051ProArg: 2.051 ± 0.453
1.436ProSer: 1.436 ± 0.515
0.615ProThr: 0.615 ± 0.176
2.667ProVal: 2.667 ± 0.372
0.205ProTrp: 0.205 ± 0.225
0.41ProTyr: 0.41 ± 0.179
0.0ProXaa: 0.0 ± 0.0
Gln
0.41GlnAla: 0.41 ± 0.179
1.026GlnCys: 1.026 ± 0.304
1.026GlnAsp: 1.026 ± 0.605
1.846GlnGlu: 1.846 ± 0.78
0.821GlnPhe: 0.821 ± 0.229
1.846GlnGly: 1.846 ± 0.854
0.0GlnHis: 0.0 ± 0.0
1.436GlnIle: 1.436 ± 0.299
3.282GlnLys: 3.282 ± 1.011
2.462GlnLeu: 2.462 ± 0.47
0.41GlnMet: 0.41 ± 0.178
1.641GlnAsn: 1.641 ± 0.351
0.205GlnPro: 0.205 ± 0.318
1.026GlnGln: 1.026 ± 0.331
3.077GlnArg: 3.077 ± 0.722
2.256GlnSer: 2.256 ± 0.524
1.846GlnThr: 1.846 ± 0.949
1.436GlnVal: 1.436 ± 0.296
0.41GlnTrp: 0.41 ± 0.179
2.051GlnTyr: 2.051 ± 0.424
0.205GlnXaa: 0.205 ± 0.141
Arg
5.744ArgAla: 5.744 ± 0.722
0.821ArgCys: 0.821 ± 0.199
3.897ArgAsp: 3.897 ± 0.98
4.923ArgGlu: 4.923 ± 0.914
3.282ArgPhe: 3.282 ± 0.978
2.051ArgGly: 2.051 ± 0.348
1.436ArgHis: 1.436 ± 0.461
3.282ArgIle: 3.282 ± 0.685
7.385ArgLys: 7.385 ± 1.195
6.564ArgLeu: 6.564 ± 1.198
2.256ArgMet: 2.256 ± 0.405
1.436ArgAsn: 1.436 ± 0.347
1.026ArgPro: 1.026 ± 0.714
1.231ArgGln: 1.231 ± 0.887
3.897ArgArg: 3.897 ± 1.613
4.103ArgSer: 4.103 ± 0.656
3.077ArgThr: 3.077 ± 0.717
3.897ArgVal: 3.897 ± 0.453
1.641ArgTrp: 1.641 ± 0.363
2.462ArgTyr: 2.462 ± 1.203
0.0ArgXaa: 0.0 ± 0.0
Ser
4.513SerAla: 4.513 ± 0.884
1.231SerCys: 1.231 ± 0.279
5.744SerAsp: 5.744 ± 0.761
6.359SerGlu: 6.359 ± 0.892
2.256SerPhe: 2.256 ± 0.419
3.897SerGly: 3.897 ± 0.679
0.821SerHis: 0.821 ± 0.363
3.487SerIle: 3.487 ± 0.839
4.923SerLys: 4.923 ± 0.539
6.564SerLeu: 6.564 ± 0.597
1.026SerMet: 1.026 ± 0.577
3.282SerAsn: 3.282 ± 0.882
1.641SerPro: 1.641 ± 0.589
2.462SerGln: 2.462 ± 0.446
4.718SerArg: 4.718 ± 1.12
6.359SerSer: 6.359 ± 2.971
5.333SerThr: 5.333 ± 1.137
7.59SerVal: 7.59 ± 1.887
1.026SerTrp: 1.026 ± 0.383
2.462SerTyr: 2.462 ± 0.825
0.0SerXaa: 0.0 ± 0.0
Thr
3.897ThrAla: 3.897 ± 0.835
0.821ThrCys: 0.821 ± 0.421
3.897ThrAsp: 3.897 ± 0.619
4.103ThrGlu: 4.103 ± 0.916
3.282ThrPhe: 3.282 ± 0.636
1.231ThrGly: 1.231 ± 0.494
1.641ThrHis: 1.641 ± 0.381
2.462ThrIle: 2.462 ± 0.649
2.256ThrLys: 2.256 ± 0.585
4.308ThrLeu: 4.308 ± 0.513
1.026ThrMet: 1.026 ± 0.274
4.718ThrAsn: 4.718 ± 0.59
1.436ThrPro: 1.436 ± 0.348
2.051ThrGln: 2.051 ± 0.453
3.077ThrArg: 3.077 ± 0.672
3.897ThrSer: 3.897 ± 0.406
2.462ThrThr: 2.462 ± 0.382
6.564ThrVal: 6.564 ± 1.154
0.205ThrTrp: 0.205 ± 0.141
1.436ThrTyr: 1.436 ± 0.646
0.0ThrXaa: 0.0 ± 0.0
Val
3.487ValAla: 3.487 ± 1.662
0.821ValCys: 0.821 ± 0.427
8.205ValAsp: 8.205 ± 0.959
6.564ValGlu: 6.564 ± 1.005
3.487ValPhe: 3.487 ± 0.738
4.923ValGly: 4.923 ± 0.539
2.051ValHis: 2.051 ± 0.695
5.538ValIle: 5.538 ± 0.898
4.923ValLys: 4.923 ± 1.142
4.718ValLeu: 4.718 ± 0.533
1.231ValMet: 1.231 ± 0.53
3.077ValAsn: 3.077 ± 0.457
1.436ValPro: 1.436 ± 0.489
1.846ValGln: 1.846 ± 0.679
5.333ValArg: 5.333 ± 0.181
5.333ValSer: 5.333 ± 1.146
2.051ValThr: 2.051 ± 0.591
5.949ValVal: 5.949 ± 1.014
0.0ValTrp: 0.0 ± 0.0
3.692ValTyr: 3.692 ± 1.123
0.0ValXaa: 0.0 ± 0.0
Trp
1.231TrpAla: 1.231 ± 0.536
0.0TrpCys: 0.0 ± 0.0
0.205TrpAsp: 0.205 ± 0.141
0.205TrpGlu: 0.205 ± 0.141
0.41TrpPhe: 0.41 ± 0.179
0.41TrpGly: 0.41 ± 0.179
0.41TrpHis: 0.41 ± 0.179
0.0TrpIle: 0.0 ± 0.0
0.41TrpLys: 0.41 ± 0.179
0.615TrpLeu: 0.615 ± 0.277
0.615TrpMet: 0.615 ± 0.259
1.026TrpAsn: 1.026 ± 0.248
0.205TrpPro: 0.205 ± 0.332
0.615TrpGln: 0.615 ± 0.627
0.41TrpArg: 0.41 ± 0.329
1.026TrpSer: 1.026 ± 0.429
0.0TrpThr: 0.0 ± 0.0
1.436TrpVal: 1.436 ± 0.639
0.615TrpTrp: 0.615 ± 0.265
0.41TrpTyr: 0.41 ± 0.179
0.0TrpXaa: 0.0 ± 0.0
Tyr
3.077TyrAla: 3.077 ± 0.742
0.0TyrCys: 0.0 ± 0.0
3.487TyrAsp: 3.487 ± 0.479
2.256TyrGlu: 2.256 ± 0.576
1.641TyrPhe: 1.641 ± 0.276
1.641TyrGly: 1.641 ± 0.257
0.615TyrHis: 0.615 ± 0.265
0.821TyrIle: 0.821 ± 0.363
1.641TyrLys: 1.641 ± 0.533
4.103TyrLeu: 4.103 ± 1.049
0.821TyrMet: 0.821 ± 0.363
1.641TyrAsn: 1.641 ± 0.704
1.436TyrPro: 1.436 ± 0.469
0.205TyrGln: 0.205 ± 0.141
2.872TyrArg: 2.872 ± 0.376
2.872TyrSer: 2.872 ± 0.584
0.821TyrThr: 0.821 ± 0.572
3.282TyrVal: 3.282 ± 0.539
0.615TyrTrp: 0.615 ± 0.328
0.615TyrTyr: 0.615 ± 0.265
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.205XaaGln: 0.205 ± 0.225
0.205XaaArg: 0.205 ± 0.141
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (4876 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski