Amino acid dipepetide frequency for Beet virus Q

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
7.28AlaAla: 7.28 ± 2.308
0.416AlaCys: 0.416 ± 0.144
4.16AlaAsp: 4.16 ± 0.961
4.576AlaGlu: 4.576 ± 0.638
2.704AlaPhe: 2.704 ± 0.311
3.744AlaGly: 3.744 ± 0.526
1.04AlaHis: 1.04 ± 0.326
2.912AlaIle: 2.912 ± 0.785
2.912AlaLys: 2.912 ± 0.904
5.408AlaLeu: 5.408 ± 1.741
2.704AlaMet: 2.704 ± 0.613
3.952AlaAsn: 3.952 ± 0.624
0.832AlaPro: 0.832 ± 0.331
2.288AlaGln: 2.288 ± 0.563
3.952AlaArg: 3.952 ± 0.759
3.952AlaSer: 3.952 ± 1.399
5.408AlaThr: 5.408 ± 0.65
4.992AlaVal: 4.992 ± 0.672
0.208AlaTrp: 0.208 ± 0.29
1.872AlaTyr: 1.872 ± 0.796
0.0AlaXaa: 0.0 ± 0.0
Cys
1.664CysAla: 1.664 ± 0.467
1.456CysCys: 1.456 ± 0.776
2.288CysAsp: 2.288 ± 0.657
2.496CysGlu: 2.496 ± 0.865
0.832CysPhe: 0.832 ± 0.299
2.288CysGly: 2.288 ± 0.591
0.208CysHis: 0.208 ± 0.279
0.624CysIle: 0.624 ± 0.224
1.872CysLys: 1.872 ± 0.346
0.832CysLeu: 0.832 ± 0.518
0.208CysMet: 0.208 ± 0.264
0.832CysAsn: 0.832 ± 0.267
0.416CysPro: 0.416 ± 0.144
2.08CysGln: 2.08 ± 0.612
0.832CysArg: 0.832 ± 0.626
1.248CysSer: 1.248 ± 0.401
0.832CysThr: 0.832 ± 0.377
1.872CysVal: 1.872 ± 0.794
0.624CysTrp: 0.624 ± 0.233
1.248CysTyr: 1.248 ± 0.47
0.0CysXaa: 0.0 ± 0.0
Asp
4.576AspAla: 4.576 ± 0.87
1.04AspCys: 1.04 ± 0.376
5.616AspAsp: 5.616 ± 1.287
4.368AspGlu: 4.368 ± 0.699
2.288AspPhe: 2.288 ± 0.427
2.704AspGly: 2.704 ± 0.392
1.872AspHis: 1.872 ± 0.632
3.12AspIle: 3.12 ± 0.998
3.536AspLys: 3.536 ± 0.487
5.408AspLeu: 5.408 ± 0.495
1.664AspMet: 1.664 ± 0.58
2.08AspAsn: 2.08 ± 0.337
2.912AspPro: 2.912 ± 0.66
2.496AspGln: 2.496 ± 0.556
2.912AspArg: 2.912 ± 0.658
8.319AspSer: 8.319 ± 0.899
3.12AspThr: 3.12 ± 0.556
6.24AspVal: 6.24 ± 1.427
0.624AspTrp: 0.624 ± 0.24
2.288AspTyr: 2.288 ± 0.333
0.0AspXaa: 0.0 ± 0.0
Glu
5.616GluAla: 5.616 ± 1.043
2.912GluCys: 2.912 ± 0.771
3.952GluAsp: 3.952 ± 0.585
4.16GluGlu: 4.16 ± 0.777
2.704GluPhe: 2.704 ± 0.914
3.744GluGly: 3.744 ± 0.526
3.328GluHis: 3.328 ± 1.638
4.784GluIle: 4.784 ± 1.013
6.24GluLys: 6.24 ± 1.367
7.28GluLeu: 7.28 ± 0.863
2.08GluMet: 2.08 ± 0.221
2.912GluAsn: 2.912 ± 1.014
1.248GluPro: 1.248 ± 0.251
1.04GluGln: 1.04 ± 0.585
4.576GluArg: 4.576 ± 0.834
5.408GluSer: 5.408 ± 0.38
4.576GluThr: 4.576 ± 0.873
3.952GluVal: 3.952 ± 0.392
0.624GluTrp: 0.624 ± 0.211
1.664GluTyr: 1.664 ± 0.436
0.0GluXaa: 0.0 ± 0.0
Phe
2.496PheAla: 2.496 ± 0.627
1.664PheCys: 1.664 ± 0.777
2.496PheAsp: 2.496 ± 0.665
2.912PheGlu: 2.912 ± 0.354
1.456PhePhe: 1.456 ± 0.264
4.16PheGly: 4.16 ± 1.323
1.456PheHis: 1.456 ± 0.431
1.664PheIle: 1.664 ± 0.336
1.248PheLys: 1.248 ± 0.421
4.784PheLeu: 4.784 ± 0.421
1.248PheMet: 1.248 ± 0.481
1.248PheAsn: 1.248 ± 0.481
1.872PhePro: 1.872 ± 0.538
0.416PheGln: 0.416 ± 0.359
1.04PheArg: 1.04 ± 0.24
5.408PheSer: 5.408 ± 0.788
2.704PheThr: 2.704 ± 0.392
3.952PheVal: 3.952 ± 0.515
0.0PheTrp: 0.0 ± 0.0
1.456PheTyr: 1.456 ± 0.46
0.0PheXaa: 0.0 ± 0.0
Gly
2.912GlyAla: 2.912 ± 0.475
1.456GlyCys: 1.456 ± 0.386
3.536GlyAsp: 3.536 ± 0.726
5.616GlyGlu: 5.616 ± 1.041
1.872GlyPhe: 1.872 ± 0.569
5.2GlyGly: 5.2 ± 0.869
1.248GlyHis: 1.248 ± 0.235
2.496GlyIle: 2.496 ± 0.581
10.191GlyLys: 10.191 ± 1.65
5.408GlyLeu: 5.408 ± 0.379
0.832GlyMet: 0.832 ± 0.288
2.912GlyAsn: 2.912 ± 0.771
1.04GlyPro: 1.04 ± 0.326
0.416GlyGln: 0.416 ± 0.347
2.704GlyArg: 2.704 ± 0.387
4.368GlySer: 4.368 ± 2.018
3.328GlyThr: 3.328 ± 0.596
3.536GlyVal: 3.536 ± 0.52
0.208GlyTrp: 0.208 ± 0.29
1.872GlyTyr: 1.872 ± 0.647
0.0GlyXaa: 0.0 ± 0.0
His
1.664HisAla: 1.664 ± 0.327
1.04HisCys: 1.04 ± 0.327
2.288HisAsp: 2.288 ± 0.575
1.248HisGlu: 1.248 ± 0.629
2.288HisPhe: 2.288 ± 0.627
1.456HisGly: 1.456 ± 0.44
0.624HisHis: 0.624 ± 0.264
1.248HisIle: 1.248 ± 0.481
1.456HisLys: 1.456 ± 0.737
1.872HisLeu: 1.872 ± 0.367
0.416HisMet: 0.416 ± 0.144
0.416HisAsn: 0.416 ± 0.379
0.208HisPro: 0.208 ± 0.234
0.832HisGln: 0.832 ± 0.368
1.04HisArg: 1.04 ± 0.275
2.704HisSer: 2.704 ± 0.681
0.416HisThr: 0.416 ± 0.144
1.664HisVal: 1.664 ± 0.604
0.416HisTrp: 0.416 ± 0.461
0.832HisTyr: 0.832 ± 0.288
0.0HisXaa: 0.0 ± 0.0
Ile
3.328IleAla: 3.328 ± 1.123
0.624IleCys: 0.624 ± 0.211
4.784IleAsp: 4.784 ± 0.762
3.536IleGlu: 3.536 ± 0.625
2.704IlePhe: 2.704 ± 0.71
1.248IleGly: 1.248 ± 0.378
1.872IleHis: 1.872 ± 0.449
3.328IleIle: 3.328 ± 0.944
2.496IleLys: 2.496 ± 0.523
4.368IleLeu: 4.368 ± 0.699
1.664IleMet: 1.664 ± 0.343
2.08IleAsn: 2.08 ± 0.798
2.288IlePro: 2.288 ± 0.789
0.208IleGln: 0.208 ± 0.234
2.912IleArg: 2.912 ± 0.426
1.456IleSer: 1.456 ± 0.442
2.288IleThr: 2.288 ± 0.617
4.16IleVal: 4.16 ± 0.357
0.0IleTrp: 0.0 ± 0.0
3.328IleTyr: 3.328 ± 0.678
0.0IleXaa: 0.0 ± 0.0
Lys
2.496LysAla: 2.496 ± 1.295
1.664LysCys: 1.664 ± 0.417
5.616LysAsp: 5.616 ± 0.54
6.656LysGlu: 6.656 ± 1.497
2.912LysPhe: 2.912 ± 0.854
3.744LysGly: 3.744 ± 1.096
1.248LysHis: 1.248 ± 0.313
2.704LysIle: 2.704 ± 0.525
3.744LysLys: 3.744 ± 1.294
6.032LysLeu: 6.032 ± 1.667
1.04LysMet: 1.04 ± 0.37
2.08LysAsn: 2.08 ± 0.819
1.872LysPro: 1.872 ± 0.448
2.496LysGln: 2.496 ± 0.443
5.616LysArg: 5.616 ± 0.718
6.24LysSer: 6.24 ± 0.435
3.744LysThr: 3.744 ± 0.954
4.16LysVal: 4.16 ± 1.167
1.664LysTrp: 1.664 ± 0.327
2.496LysTyr: 2.496 ± 0.477
0.0LysXaa: 0.0 ± 0.0
Leu
3.744LeuAla: 3.744 ± 1.23
1.872LeuCys: 1.872 ± 0.508
5.408LeuAsp: 5.408 ± 1.104
4.784LeuGlu: 4.784 ± 0.859
4.992LeuPhe: 4.992 ± 0.605
4.16LeuGly: 4.16 ± 1.242
2.288LeuHis: 2.288 ± 0.789
2.912LeuIle: 2.912 ± 0.793
6.032LeuLys: 6.032 ± 0.442
9.567LeuLeu: 9.567 ± 1.788
2.496LeuMet: 2.496 ± 0.556
4.368LeuAsn: 4.368 ± 0.515
3.744LeuPro: 3.744 ± 0.465
3.536LeuGln: 3.536 ± 0.273
4.368LeuArg: 4.368 ± 1.428
6.24LeuSer: 6.24 ± 2.127
6.448LeuThr: 6.448 ± 0.881
8.527LeuVal: 8.527 ± 0.91
1.248LeuTrp: 1.248 ± 0.268
2.496LeuTyr: 2.496 ± 0.613
0.0LeuXaa: 0.0 ± 0.0
Met
2.912MetAla: 2.912 ± 1.014
1.04MetCys: 1.04 ± 0.316
1.248MetAsp: 1.248 ± 0.298
1.456MetGlu: 1.456 ± 0.386
1.456MetPhe: 1.456 ± 0.426
0.832MetGly: 0.832 ± 0.288
0.0MetHis: 0.0 ± 0.0
2.704MetIle: 2.704 ± 0.975
2.704MetLys: 2.704 ± 0.597
2.08MetLeu: 2.08 ± 0.516
0.832MetMet: 0.832 ± 0.288
1.04MetAsn: 1.04 ± 0.37
0.624MetPro: 0.624 ± 0.408
0.624MetGln: 0.624 ± 0.224
1.872MetArg: 1.872 ± 0.564
2.496MetSer: 2.496 ± 0.834
1.04MetThr: 1.04 ± 0.37
1.248MetVal: 1.248 ± 0.569
0.0MetTrp: 0.0 ± 0.0
0.208MetTyr: 0.208 ± 0.143
0.0MetXaa: 0.0 ± 0.0
Asn
1.872AsnAla: 1.872 ± 0.531
1.664AsnCys: 1.664 ± 0.355
2.704AsnAsp: 2.704 ± 0.678
3.328AsnGlu: 3.328 ± 0.475
1.872AsnPhe: 1.872 ± 0.601
4.576AsnGly: 4.576 ± 0.925
1.04AsnHis: 1.04 ± 0.275
1.456AsnIle: 1.456 ± 0.856
3.744AsnLys: 3.744 ± 0.776
2.496AsnLeu: 2.496 ± 0.951
0.624AsnMet: 0.624 ± 0.24
0.832AsnAsn: 0.832 ± 0.251
0.416AsnPro: 0.416 ± 0.379
1.04AsnGln: 1.04 ± 0.27
1.872AsnArg: 1.872 ± 0.578
2.912AsnSer: 2.912 ± 0.953
1.248AsnThr: 1.248 ± 0.517
2.704AsnVal: 2.704 ± 0.456
0.832AsnTrp: 0.832 ± 0.368
1.248AsnTyr: 1.248 ± 0.317
0.0AsnXaa: 0.0 ± 0.0
Pro
2.704ProAla: 2.704 ± 0.678
1.04ProCys: 1.04 ± 0.667
1.872ProAsp: 1.872 ± 0.706
1.04ProGlu: 1.04 ± 0.258
1.248ProPhe: 1.248 ± 0.27
2.496ProGly: 2.496 ± 0.399
0.208ProHis: 0.208 ± 0.143
2.08ProIle: 2.08 ± 0.522
2.704ProLys: 2.704 ± 0.551
2.496ProLeu: 2.496 ± 0.635
1.248ProMet: 1.248 ± 0.317
1.248ProAsn: 1.248 ± 0.378
0.832ProPro: 0.832 ± 0.518
1.248ProGln: 1.248 ± 0.273
3.328ProArg: 3.328 ± 0.465
1.248ProSer: 1.248 ± 0.454
1.248ProThr: 1.248 ± 0.401
2.08ProVal: 2.08 ± 0.762
0.0ProTrp: 0.0 ± 0.0
0.624ProTyr: 0.624 ± 0.24
0.0ProXaa: 0.0 ± 0.0
Gln
0.832GlnAla: 0.832 ± 0.218
0.416GlnCys: 0.416 ± 0.255
1.04GlnAsp: 1.04 ± 0.979
2.496GlnGlu: 2.496 ± 0.29
1.04GlnPhe: 1.04 ± 0.465
3.536GlnGly: 3.536 ± 0.445
0.208GlnHis: 0.208 ± 0.273
1.664GlnIle: 1.664 ± 0.607
2.912GlnLys: 2.912 ± 0.851
3.328GlnLeu: 3.328 ± 0.594
0.208GlnMet: 0.208 ± 0.142
0.832GlnAsn: 0.832 ± 0.266
0.624GlnPro: 0.624 ± 0.427
0.624GlnGln: 0.624 ± 0.24
2.08GlnArg: 2.08 ± 0.551
2.704GlnSer: 2.704 ± 0.796
2.08GlnThr: 2.08 ± 0.416
1.04GlnVal: 1.04 ± 0.24
0.416GlnTrp: 0.416 ± 0.144
0.832GlnTyr: 0.832 ± 0.34
0.208GlnXaa: 0.208 ± 0.143
Arg
3.12ArgAla: 3.12 ± 0.649
0.832ArgCys: 0.832 ± 0.288
3.12ArgAsp: 3.12 ± 0.804
2.704ArgGlu: 2.704 ± 0.71
3.536ArgPhe: 3.536 ± 0.432
3.952ArgGly: 3.952 ± 0.781
1.456ArgHis: 1.456 ± 0.905
3.952ArgIle: 3.952 ± 0.639
3.744ArgLys: 3.744 ± 0.685
5.408ArgLeu: 5.408 ± 1.747
2.08ArgMet: 2.08 ± 0.567
2.496ArgAsn: 2.496 ± 0.444
1.664ArgPro: 1.664 ± 0.363
1.664ArgGln: 1.664 ± 0.205
3.744ArgArg: 3.744 ± 1.105
3.952ArgSer: 3.952 ± 0.689
4.576ArgThr: 4.576 ± 0.488
4.16ArgVal: 4.16 ± 1.058
0.624ArgTrp: 0.624 ± 0.24
2.288ArgTyr: 2.288 ± 0.748
0.0ArgXaa: 0.0 ± 0.0
Ser
4.784SerAla: 4.784 ± 1.405
1.456SerCys: 1.456 ± 0.589
7.488SerAsp: 7.488 ± 1.239
6.032SerGlu: 6.032 ± 0.747
2.496SerPhe: 2.496 ± 0.295
2.704SerGly: 2.704 ± 1.388
2.496SerHis: 2.496 ± 0.778
2.912SerIle: 2.912 ± 0.702
4.16SerLys: 4.16 ± 1.119
6.24SerLeu: 6.24 ± 0.722
1.872SerMet: 1.872 ± 0.546
3.12SerAsn: 3.12 ± 1.292
2.912SerPro: 2.912 ± 0.79
1.872SerGln: 1.872 ± 0.607
3.952SerArg: 3.952 ± 0.784
7.696SerSer: 7.696 ± 1.118
3.952SerThr: 3.952 ± 0.598
7.488SerVal: 7.488 ± 1.76
0.624SerTrp: 0.624 ± 0.264
1.872SerTyr: 1.872 ± 1.479
0.208SerXaa: 0.208 ± 0.29
Thr
3.328ThrAla: 3.328 ± 0.811
0.832ThrCys: 0.832 ± 0.54
1.456ThrAsp: 1.456 ± 0.402
5.2ThrGlu: 5.2 ± 1.475
2.912ThrPhe: 2.912 ± 0.728
4.576ThrGly: 4.576 ± 0.717
1.456ThrHis: 1.456 ± 0.386
2.288ThrIle: 2.288 ± 0.431
2.08ThrLys: 2.08 ± 0.68
5.616ThrLeu: 5.616 ± 0.893
2.288ThrMet: 2.288 ± 0.575
1.664ThrAsn: 1.664 ± 0.298
1.456ThrPro: 1.456 ± 0.298
2.288ThrGln: 2.288 ± 1.292
4.16ThrArg: 4.16 ± 0.668
2.912ThrSer: 2.912 ± 0.361
1.872ThrThr: 1.872 ± 0.338
5.2ThrVal: 5.2 ± 0.487
0.624ThrTrp: 0.624 ± 0.24
1.872ThrTyr: 1.872 ± 0.4
0.0ThrXaa: 0.0 ± 0.0
Val
6.24ValAla: 6.24 ± 1.218
2.496ValCys: 2.496 ± 0.564
6.448ValAsp: 6.448 ± 0.69
5.824ValGlu: 5.824 ± 0.617
3.536ValPhe: 3.536 ± 0.75
2.912ValGly: 2.912 ± 0.539
2.08ValHis: 2.08 ± 0.329
4.992ValIle: 4.992 ± 0.765
5.616ValLys: 5.616 ± 1.119
5.408ValLeu: 5.408 ± 0.84
1.456ValMet: 1.456 ± 0.501
2.912ValAsn: 2.912 ± 0.567
4.784ValPro: 4.784 ± 0.567
2.08ValGln: 2.08 ± 0.457
4.368ValArg: 4.368 ± 1.354
3.952ValSer: 3.952 ± 1.376
2.496ValThr: 2.496 ± 0.311
6.448ValVal: 6.448 ± 1.3
0.832ValTrp: 0.832 ± 0.326
1.872ValTyr: 1.872 ± 0.445
0.0ValXaa: 0.0 ± 0.0
Trp
1.248TrpAla: 1.248 ± 0.433
0.0TrpCys: 0.0 ± 0.0
0.0TrpAsp: 0.0 ± 0.0
0.624TrpGlu: 0.624 ± 0.24
0.0TrpPhe: 0.0 ± 0.0
0.832TrpGly: 0.832 ± 0.218
0.0TrpHis: 0.0 ± 0.0
0.0TrpIle: 0.0 ± 0.0
0.624TrpLys: 0.624 ± 0.211
1.456TrpLeu: 1.456 ± 0.327
0.624TrpMet: 0.624 ± 0.232
0.208TrpAsn: 0.208 ± 0.143
0.208TrpPro: 0.208 ± 0.279
0.416TrpGln: 0.416 ± 0.407
0.416TrpArg: 0.416 ± 0.461
1.04TrpSer: 1.04 ± 0.336
0.832TrpThr: 0.832 ± 0.922
0.416TrpVal: 0.416 ± 0.379
0.208TrpTrp: 0.208 ± 0.143
0.832TrpTyr: 0.832 ± 0.288
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.496TyrAla: 2.496 ± 0.789
0.832TyrCys: 0.832 ± 0.326
1.664TyrAsp: 1.664 ± 0.371
3.744TyrGlu: 3.744 ± 0.56
0.832TyrPhe: 0.832 ± 0.326
2.288TyrGly: 2.288 ± 0.55
0.0TyrHis: 0.0 ± 0.0
0.832TyrIle: 0.832 ± 0.288
0.208TyrLys: 0.208 ± 0.143
3.744TyrLeu: 3.744 ± 0.436
0.624TyrMet: 0.624 ± 0.212
1.248TyrAsn: 1.248 ± 0.586
1.04TyrPro: 1.04 ± 0.325
1.456TyrGln: 1.456 ± 0.416
3.12TyrArg: 3.12 ± 0.576
2.08TyrSer: 2.08 ± 0.359
1.872TyrThr: 1.872 ± 0.632
2.912TyrVal: 2.912 ± 0.612
0.208TyrTrp: 0.208 ± 0.279
2.08TyrTyr: 2.08 ± 0.667
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.208XaaGln: 0.208 ± 0.29
0.208XaaArg: 0.208 ± 0.143
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 9 proteins (4809 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski