Amino acid dipepetide frequency for Tortoise microvirus 56

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
2.915AlaAla: 2.915 ± 1.339
1.166AlaCys: 1.166 ± 1.18
1.749AlaAsp: 1.749 ± 0.702
4.665AlaGlu: 4.665 ± 1.899
1.749AlaPhe: 1.749 ± 0.797
4.665AlaGly: 4.665 ± 1.67
2.332AlaHis: 2.332 ± 1.124
2.915AlaIle: 2.915 ± 1.225
1.749AlaLys: 1.749 ± 1.271
5.831AlaLeu: 5.831 ± 1.827
0.583AlaMet: 0.583 ± 0.733
1.166AlaAsn: 1.166 ± 0.562
1.166AlaPro: 1.166 ± 0.847
5.831AlaGln: 5.831 ± 2.349
1.749AlaArg: 1.749 ± 0.946
1.749AlaSer: 1.749 ± 1.162
4.082AlaThr: 4.082 ± 0.785
3.499AlaVal: 3.499 ± 1.784
0.0AlaTrp: 0.0 ± 0.0
1.749AlaTyr: 1.749 ± 0.877
0.0AlaXaa: 0.0 ± 0.0
Cys
0.0CysAla: 0.0 ± 0.0
1.166CysCys: 1.166 ± 0.941
1.166CysAsp: 1.166 ± 0.626
0.583CysGlu: 0.583 ± 0.423
1.749CysPhe: 1.749 ± 0.99
0.0CysGly: 0.0 ± 0.0
0.583CysHis: 0.583 ± 0.423
0.0CysIle: 0.0 ± 0.0
0.0CysLys: 0.0 ± 0.0
2.915CysLeu: 2.915 ± 1.809
0.0CysMet: 0.0 ± 0.0
0.0CysAsn: 0.0 ± 0.0
1.749CysPro: 1.749 ± 1.336
0.0CysGln: 0.0 ± 0.0
1.166CysArg: 1.166 ± 0.559
1.166CysSer: 1.166 ± 0.559
0.583CysThr: 0.583 ± 0.668
1.749CysVal: 1.749 ± 1.069
0.0CysTrp: 0.0 ± 0.0
0.583CysTyr: 0.583 ± 0.733
0.0CysXaa: 0.0 ± 0.0
Asp
4.082AspAla: 4.082 ± 2.059
1.166AspCys: 1.166 ± 0.967
3.499AspAsp: 3.499 ± 1.639
2.915AspGlu: 2.915 ± 0.905
2.332AspPhe: 2.332 ± 1.298
1.166AspGly: 1.166 ± 0.847
0.0AspHis: 0.0 ± 0.0
1.749AspIle: 1.749 ± 0.978
1.749AspLys: 1.749 ± 0.892
4.082AspLeu: 4.082 ± 1.241
2.915AspMet: 2.915 ± 1.758
2.332AspAsn: 2.332 ± 0.884
1.166AspPro: 1.166 ± 0.847
1.749AspGln: 1.749 ± 0.718
1.749AspArg: 1.749 ± 0.718
5.248AspSer: 5.248 ± 0.985
3.499AspThr: 3.499 ± 1.1
5.831AspVal: 5.831 ± 2.573
0.583AspTrp: 0.583 ± 0.668
4.665AspTyr: 4.665 ± 2.149
0.0AspXaa: 0.0 ± 0.0
Glu
2.915GluAla: 2.915 ± 1.58
0.583GluCys: 0.583 ± 0.733
1.166GluAsp: 1.166 ± 0.626
1.749GluGlu: 1.749 ± 0.819
2.332GluPhe: 2.332 ± 1.512
1.166GluGly: 1.166 ± 0.967
1.166GluHis: 1.166 ± 0.626
2.915GluIle: 2.915 ± 1.337
2.915GluLys: 2.915 ± 1.481
4.082GluLeu: 4.082 ± 1.802
0.583GluMet: 0.583 ± 0.549
3.499GluAsn: 3.499 ± 2.694
0.583GluPro: 0.583 ± 0.641
3.499GluGln: 3.499 ± 2.312
2.332GluArg: 2.332 ± 1.676
1.749GluSer: 1.749 ± 1.045
4.082GluThr: 4.082 ± 0.757
1.749GluVal: 1.749 ± 0.9
0.0GluTrp: 0.0 ± 0.0
3.499GluTyr: 3.499 ± 1.638
0.0GluXaa: 0.0 ± 0.0
Phe
1.166PheAla: 1.166 ± 0.562
1.166PheCys: 1.166 ± 0.941
4.082PheAsp: 4.082 ± 1.877
2.332PheGlu: 2.332 ± 1.124
2.332PhePhe: 2.332 ± 1.045
2.915PheGly: 2.915 ± 1.567
1.166PheHis: 1.166 ± 0.685
5.248PheIle: 5.248 ± 3.865
4.082PheLys: 4.082 ± 1.335
4.665PheLeu: 4.665 ± 4.602
0.0PheMet: 0.0 ± 0.0
2.915PheAsn: 2.915 ± 1.005
0.583PhePro: 0.583 ± 0.423
1.166PheGln: 1.166 ± 0.562
2.332PheArg: 2.332 ± 0.59
3.499PheSer: 3.499 ± 2.101
3.499PheThr: 3.499 ± 1.12
2.915PheVal: 2.915 ± 1.393
0.0PheTrp: 0.0 ± 0.0
9.913PheTyr: 9.913 ± 7.272
0.0PheXaa: 0.0 ± 0.0
Gly
1.166GlyAla: 1.166 ± 0.562
0.0GlyCys: 0.0 ± 0.0
1.749GlyAsp: 1.749 ± 0.818
2.915GlyGlu: 2.915 ± 1.081
1.749GlyPhe: 1.749 ± 1.045
2.915GlyGly: 2.915 ± 1.58
0.0GlyHis: 0.0 ± 0.0
1.166GlyIle: 1.166 ± 0.847
1.166GlyLys: 1.166 ± 0.847
4.082GlyLeu: 4.082 ± 1.409
1.749GlyMet: 1.749 ± 1.27
2.332GlyAsn: 2.332 ± 0.671
0.583GlyPro: 0.583 ± 0.667
3.499GlyGln: 3.499 ± 1.638
2.915GlyArg: 2.915 ± 2.149
5.831GlySer: 5.831 ± 1.955
4.082GlyThr: 4.082 ± 1.892
2.915GlyVal: 2.915 ± 1.567
0.583GlyTrp: 0.583 ± 0.641
3.499GlyTyr: 3.499 ± 1.444
0.0GlyXaa: 0.0 ± 0.0
His
1.749HisAla: 1.749 ± 1.045
1.166HisCys: 1.166 ± 0.559
0.583HisAsp: 0.583 ± 0.423
0.0HisGlu: 0.0 ± 0.0
2.332HisPhe: 2.332 ± 1.693
1.166HisGly: 1.166 ± 0.847
0.583HisHis: 0.583 ± 0.423
1.749HisIle: 1.749 ± 1.42
0.0HisLys: 0.0 ± 0.0
0.583HisLeu: 0.583 ± 0.59
0.0HisMet: 0.0 ± 0.0
1.749HisAsn: 1.749 ± 1.045
0.583HisPro: 0.583 ± 0.423
0.0HisGln: 0.0 ± 0.0
0.583HisArg: 0.583 ± 0.423
0.583HisSer: 0.583 ± 0.668
0.0HisThr: 0.0 ± 0.0
1.749HisVal: 1.749 ± 1.112
0.0HisTrp: 0.0 ± 0.0
1.166HisTyr: 1.166 ± 0.847
0.0HisXaa: 0.0 ± 0.0
Ile
5.248IleAla: 5.248 ± 1.888
1.166IleCys: 1.166 ± 0.559
3.499IleAsp: 3.499 ± 1.104
2.915IleGlu: 2.915 ± 1.13
1.166IlePhe: 1.166 ± 0.847
4.665IleGly: 4.665 ± 1.791
0.583IleHis: 0.583 ± 0.668
10.496IleIle: 10.496 ± 6.106
3.499IleLys: 3.499 ± 1.361
8.746IleLeu: 8.746 ± 4.605
2.332IleMet: 2.332 ± 1.027
3.499IleAsn: 3.499 ± 1.76
4.082IlePro: 4.082 ± 1.529
2.915IleGln: 2.915 ± 1.747
5.248IleArg: 5.248 ± 3.939
5.248IleSer: 5.248 ± 2.09
4.665IleThr: 4.665 ± 1.156
4.082IleVal: 4.082 ± 3.223
0.0IleTrp: 0.0 ± 0.0
1.749IleTyr: 1.749 ± 1.422
0.0IleXaa: 0.0 ± 0.0
Lys
0.0LysAla: 0.0 ± 0.0
0.583LysCys: 0.583 ± 0.423
0.0LysAsp: 0.0 ± 0.0
1.166LysGlu: 1.166 ± 0.562
3.499LysPhe: 3.499 ± 1.167
2.915LysGly: 2.915 ± 1.192
0.583LysHis: 0.583 ± 0.59
4.665LysIle: 4.665 ± 1.166
0.583LysLys: 0.583 ± 0.668
3.499LysLeu: 3.499 ± 1.658
0.0LysMet: 0.0 ± 0.0
2.332LysAsn: 2.332 ± 1.599
2.332LysPro: 2.332 ± 1.196
1.749LysGln: 1.749 ± 1.156
3.499LysArg: 3.499 ± 1.263
5.248LysSer: 5.248 ± 1.93
2.915LysThr: 2.915 ± 0.775
2.915LysVal: 2.915 ± 0.89
0.583LysTrp: 0.583 ± 0.565
1.166LysTyr: 1.166 ± 0.772
0.0LysXaa: 0.0 ± 0.0
Leu
5.248LeuAla: 5.248 ± 1.414
1.749LeuCys: 1.749 ± 0.718
5.831LeuAsp: 5.831 ± 2.299
1.749LeuGlu: 1.749 ± 1.266
10.496LeuPhe: 10.496 ± 8.231
5.248LeuGly: 5.248 ± 2.19
1.749LeuHis: 1.749 ± 0.738
4.082LeuIle: 4.082 ± 2.915
3.499LeuLys: 3.499 ± 1.369
9.913LeuLeu: 9.913 ± 4.808
2.332LeuMet: 2.332 ± 0.955
6.414LeuAsn: 6.414 ± 1.271
2.332LeuPro: 2.332 ± 1.156
2.915LeuGln: 2.915 ± 1.616
8.163LeuArg: 8.163 ± 4.145
9.913LeuSer: 9.913 ± 3.439
7.58LeuThr: 7.58 ± 3.575
4.665LeuVal: 4.665 ± 1.756
0.583LeuTrp: 0.583 ± 0.423
2.915LeuTyr: 2.915 ± 0.99
0.0LeuXaa: 0.0 ± 0.0
Met
2.332MetAla: 2.332 ± 0.702
0.0MetCys: 0.0 ± 0.0
1.749MetAsp: 1.749 ± 1.069
2.332MetGlu: 2.332 ± 1.11
1.749MetPhe: 1.749 ± 0.962
0.583MetGly: 0.583 ± 0.565
1.166MetHis: 1.166 ± 0.756
1.749MetIle: 1.749 ± 0.642
0.583MetLys: 0.583 ± 0.59
5.248MetLeu: 5.248 ± 1.625
0.0MetMet: 0.0 ± 0.0
1.749MetAsn: 1.749 ± 0.946
1.166MetPro: 1.166 ± 0.562
2.332MetGln: 2.332 ± 0.888
1.166MetArg: 1.166 ± 0.559
0.583MetSer: 0.583 ± 0.423
0.583MetThr: 0.583 ± 0.641
0.583MetVal: 0.583 ± 0.641
0.583MetTrp: 0.583 ± 0.565
0.583MetTyr: 0.583 ± 0.668
0.0MetXaa: 0.0 ± 0.0
Asn
2.915AsnAla: 2.915 ± 1.58
1.166AsnCys: 1.166 ± 0.559
2.915AsnAsp: 2.915 ± 1.577
4.665AsnGlu: 4.665 ± 2.361
2.332AsnPhe: 2.332 ± 0.967
1.749AsnGly: 1.749 ± 0.923
1.749AsnHis: 1.749 ± 0.819
5.831AsnIle: 5.831 ± 2.14
0.583AsnLys: 0.583 ± 0.565
3.499AsnLeu: 3.499 ± 1.188
1.749AsnMet: 1.749 ± 0.791
4.665AsnAsn: 4.665 ± 2.22
2.332AsnPro: 2.332 ± 0.806
0.583AsnGln: 0.583 ± 0.641
2.915AsnArg: 2.915 ± 1.339
6.997AsnSer: 6.997 ± 2.746
2.915AsnThr: 2.915 ± 1.201
4.082AsnVal: 4.082 ± 0.933
0.583AsnTrp: 0.583 ± 0.565
4.665AsnTyr: 4.665 ± 1.178
0.0AsnXaa: 0.0 ± 0.0
Pro
3.499ProAla: 3.499 ± 1.129
0.583ProCys: 0.583 ± 0.423
1.749ProAsp: 1.749 ± 0.702
2.332ProGlu: 2.332 ± 1.252
0.583ProPhe: 0.583 ± 0.423
0.583ProGly: 0.583 ± 0.423
0.0ProHis: 0.0 ± 0.0
4.665ProIle: 4.665 ± 1.77
1.749ProLys: 1.749 ± 0.638
3.499ProLeu: 3.499 ± 1.099
0.0ProMet: 0.0 ± 0.0
1.166ProAsn: 1.166 ± 0.685
1.166ProPro: 1.166 ± 0.941
1.166ProGln: 1.166 ± 0.559
2.332ProArg: 2.332 ± 1.252
4.082ProSer: 4.082 ± 1.925
1.166ProThr: 1.166 ± 0.866
2.332ProVal: 2.332 ± 1.196
1.166ProTrp: 1.166 ± 0.626
1.166ProTyr: 1.166 ± 0.562
0.0ProXaa: 0.0 ± 0.0
Gln
2.915GlnAla: 2.915 ± 2.32
0.583GlnCys: 0.583 ± 0.668
1.749GlnAsp: 1.749 ± 0.978
1.166GlnGlu: 1.166 ± 0.562
2.332GlnPhe: 2.332 ± 0.782
0.583GlnGly: 0.583 ± 0.668
0.583GlnHis: 0.583 ± 0.423
4.665GlnIle: 4.665 ± 2.148
2.332GlnLys: 2.332 ± 0.997
4.665GlnLeu: 4.665 ± 3.254
2.332GlnMet: 2.332 ± 1.656
2.332GlnAsn: 2.332 ± 1.11
1.166GlnPro: 1.166 ± 0.847
7.58GlnGln: 7.58 ± 3.529
1.749GlnArg: 1.749 ± 1.271
5.831GlnSer: 5.831 ± 2.1
3.499GlnThr: 3.499 ± 1.861
1.166GlnVal: 1.166 ± 0.562
0.0GlnTrp: 0.0 ± 0.0
1.166GlnTyr: 1.166 ± 0.562
0.0GlnXaa: 0.0 ± 0.0
Arg
2.332ArgAla: 2.332 ± 1.071
0.0ArgCys: 0.0 ± 0.0
4.665ArgAsp: 4.665 ± 1.587
1.166ArgGlu: 1.166 ± 0.833
2.332ArgPhe: 2.332 ± 1.584
2.915ArgGly: 2.915 ± 1.0
0.0ArgHis: 0.0 ± 0.0
4.082ArgIle: 4.082 ± 1.178
3.499ArgLys: 3.499 ± 2.385
7.58ArgLeu: 7.58 ± 1.014
1.749ArgMet: 1.749 ± 1.072
4.665ArgAsn: 4.665 ± 0.878
0.583ArgPro: 0.583 ± 0.59
3.499ArgGln: 3.499 ± 2.12
2.332ArgArg: 2.332 ± 1.676
4.082ArgSer: 4.082 ± 1.318
2.915ArgThr: 2.915 ± 1.081
4.665ArgVal: 4.665 ± 1.352
1.749ArgTrp: 1.749 ± 0.819
2.915ArgTyr: 2.915 ± 1.126
0.0ArgXaa: 0.0 ± 0.0
Ser
1.749SerAla: 1.749 ± 0.638
0.583SerCys: 0.583 ± 0.59
2.332SerAsp: 2.332 ± 1.347
3.499SerGlu: 3.499 ± 2.134
4.665SerPhe: 4.665 ± 1.21
5.831SerGly: 5.831 ± 1.194
1.166SerHis: 1.166 ± 0.559
5.248SerIle: 5.248 ± 1.761
5.248SerLys: 5.248 ± 1.533
9.913SerLeu: 9.913 ± 2.701
2.915SerMet: 2.915 ± 1.281
3.499SerAsn: 3.499 ± 1.428
4.082SerPro: 4.082 ± 1.563
4.665SerGln: 4.665 ± 1.986
4.082SerArg: 4.082 ± 0.977
9.913SerSer: 9.913 ± 2.837
5.248SerThr: 5.248 ± 1.624
9.913SerVal: 9.913 ± 2.553
0.583SerTrp: 0.583 ± 0.733
4.665SerTyr: 4.665 ± 1.523
0.0SerXaa: 0.0 ± 0.0
Thr
5.248ThrAla: 5.248 ± 2.338
0.0ThrCys: 0.0 ± 0.0
2.915ThrAsp: 2.915 ± 1.795
1.166ThrGlu: 1.166 ± 0.685
1.166ThrPhe: 1.166 ± 0.728
2.332ThrGly: 2.332 ± 0.806
1.749ThrHis: 1.749 ± 0.962
2.332ThrIle: 2.332 ± 1.117
0.583ThrLys: 0.583 ± 0.668
5.248ThrLeu: 5.248 ± 2.668
2.332ThrMet: 2.332 ± 0.678
4.665ThrAsn: 4.665 ± 1.294
4.082ThrPro: 4.082 ± 1.829
2.915ThrGln: 2.915 ± 0.951
3.499ThrArg: 3.499 ± 1.358
8.746ThrSer: 8.746 ± 1.57
4.082ThrThr: 4.082 ± 1.493
4.082ThrVal: 4.082 ± 2.438
0.583ThrTrp: 0.583 ± 0.565
1.749ThrTyr: 1.749 ± 0.797
0.0ThrXaa: 0.0 ± 0.0
Val
2.915ValAla: 2.915 ± 1.747
0.583ValCys: 0.583 ± 0.59
6.414ValAsp: 6.414 ± 1.734
2.915ValGlu: 2.915 ± 1.724
2.332ValPhe: 2.332 ± 1.355
3.499ValGly: 3.499 ± 1.414
1.166ValHis: 1.166 ± 0.847
6.414ValIle: 6.414 ± 3.785
5.248ValLys: 5.248 ± 2.37
5.831ValLeu: 5.831 ± 2.363
3.499ValMet: 3.499 ± 0.559
4.082ValAsn: 4.082 ± 1.455
2.915ValPro: 2.915 ± 1.112
1.166ValGln: 1.166 ± 0.562
4.082ValArg: 4.082 ± 2.7
4.082ValSer: 4.082 ± 1.463
3.499ValThr: 3.499 ± 1.587
4.082ValVal: 4.082 ± 1.884
0.583ValTrp: 0.583 ± 0.667
2.332ValTyr: 2.332 ± 0.967
0.0ValXaa: 0.0 ± 0.0
Trp
0.583TrpAla: 0.583 ± 0.423
0.0TrpCys: 0.0 ± 0.0
0.583TrpAsp: 0.583 ± 0.565
1.166TrpGlu: 1.166 ± 1.131
0.583TrpPhe: 0.583 ± 0.423
0.0TrpGly: 0.0 ± 0.0
0.0TrpHis: 0.0 ± 0.0
1.166TrpIle: 1.166 ± 0.898
0.0TrpLys: 0.0 ± 0.0
0.0TrpLeu: 0.0 ± 0.0
0.0TrpMet: 0.0 ± 0.0
1.749TrpAsn: 1.749 ± 0.791
0.0TrpPro: 0.0 ± 0.0
0.0TrpGln: 0.0 ± 0.0
1.166TrpArg: 1.166 ± 0.833
0.0TrpSer: 0.0 ± 0.0
0.583TrpThr: 0.583 ± 0.423
0.583TrpVal: 0.583 ± 0.668
0.0TrpTrp: 0.0 ± 0.0
0.0TrpTyr: 0.0 ± 0.0
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.332TyrAla: 2.332 ± 1.162
1.749TyrCys: 1.749 ± 0.962
4.082TyrAsp: 4.082 ± 1.589
1.166TyrGlu: 1.166 ± 1.024
8.163TyrPhe: 8.163 ± 5.915
0.0TyrGly: 0.0 ± 0.0
0.0TyrHis: 0.0 ± 0.0
4.665TyrIle: 4.665 ± 1.42
1.166TyrLys: 1.166 ± 0.772
4.082TyrLeu: 4.082 ± 1.572
1.166TyrMet: 1.166 ± 0.827
4.082TyrAsn: 4.082 ± 1.455
2.332TyrPro: 2.332 ± 0.967
1.166TyrGln: 1.166 ± 0.562
4.665TyrArg: 4.665 ± 2.342
5.248TyrSer: 5.248 ± 2.332
0.0TyrThr: 0.0 ± 0.0
4.082TyrVal: 4.082 ± 1.339
0.0TyrTrp: 0.0 ± 0.0
5.831TyrTyr: 5.831 ± 4.513
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (1716 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski