Amino acid dipepetide frequency for Pseudocercospora fijiensis (strain CIRAD86) (Black leaf streak disease fungus) (Mycosphaerella fijiensis)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	9.355AlaAla: 9.355 ± 0.058	1.291AlaCys: 1.291 ± 0.017	4.357AlaAsp: 4.357 ± 0.029	5.478AlaGlu: 5.478 ± 0.036	3.359AlaPhe: 3.359 ± 0.027	5.941AlaGly: 5.941 ± 0.034	1.941AlaHis: 1.941 ± 0.021	4.347AlaIle: 4.347 ± 0.034	4.497AlaLys: 4.497 ± 0.035	7.908AlaLeu: 7.908 ± 0.044	2.08AlaMet: 2.08 ± 0.018	3.063AlaAsn: 3.063 ± 0.025	4.765AlaPro: 4.765 ± 0.04	3.711AlaGln: 3.711 ± 0.032	5.368AlaArg: 5.368 ± 0.036	7.438AlaSer: 7.438 ± 0.046	5.325AlaThr: 5.325 ± 0.032	5.367AlaVal: 5.367 ± 0.039	1.32AlaTrp: 1.32 ± 0.017	2.301AlaTyr: 2.301 ± 0.023	0.0AlaXaa: 0.0 ± 0.0
Cys	1.113CysAla: 1.113 ± 0.013	0.352CysCys: 0.352 ± 0.008	0.724CysAsp: 0.724 ± 0.013	0.713CysGlu: 0.713 ± 0.014	0.631CysPhe: 0.631 ± 0.011	1.04CysGly: 1.04 ± 0.013	0.429CysHis: 0.429 ± 0.01	0.823CysIle: 0.823 ± 0.014	0.603CysLys: 0.603 ± 0.012	1.397CysLeu: 1.397 ± 0.017	0.339CysMet: 0.339 ± 0.008	0.48CysAsn: 0.48 ± 0.01	0.739CysPro: 0.739 ± 0.014	0.512CysGln: 0.512 ± 0.011	0.956CysArg: 0.956 ± 0.013	1.144CysSer: 1.144 ± 0.016	0.839CysThr: 0.839 ± 0.014	0.855CysVal: 0.855 ± 0.014	0.259CysTrp: 0.259 ± 0.007	0.416CysTyr: 0.416 ± 0.009	0.0CysXaa: 0.0 ± 0.0
Asp	4.851AspAla: 4.851 ± 0.031	0.709AspCys: 0.709 ± 0.012	4.105AspAsp: 4.105 ± 0.046	4.362AspGlu: 4.362 ± 0.036	2.288AspPhe: 2.288 ± 0.02	4.019AspGly: 4.019 ± 0.029	1.33AspHis: 1.33 ± 0.016	2.781AspIle: 2.781 ± 0.023	2.206AspLys: 2.206 ± 0.025	5.023AspLeu: 5.023 ± 0.033	1.274AspMet: 1.274 ± 0.015	1.671AspAsn: 1.671 ± 0.022	2.991AspPro: 2.991 ± 0.026	1.994AspGln: 1.994 ± 0.021	3.106AspArg: 3.106 ± 0.026	3.915AspSer: 3.915 ± 0.029	2.873AspThr: 2.873 ± 0.022	3.656AspVal: 3.656 ± 0.031	0.905AspTrp: 0.905 ± 0.012	1.528AspTyr: 1.528 ± 0.02	0.0AspXaa: 0.0 ± 0.0
Glu	5.471GluAla: 5.471 ± 0.037	0.697GluCys: 0.697 ± 0.011	4.111GluAsp: 4.111 ± 0.033	5.248GluGlu: 5.248 ± 0.05	1.804GluPhe: 1.804 ± 0.02	3.727GluGly: 3.727 ± 0.028	1.588GluHis: 1.588 ± 0.018	2.976GluIle: 2.976 ± 0.028	3.782GluLys: 3.782 ± 0.037	5.163GluLeu: 5.163 ± 0.038	1.496GluMet: 1.496 ± 0.017	2.191GluAsn: 2.191 ± 0.02	2.631GluPro: 2.631 ± 0.025	2.733GluGln: 2.733 ± 0.027	4.188GluArg: 4.188 ± 0.036	4.042GluSer: 4.042 ± 0.026	3.204GluThr: 3.204 ± 0.026	3.528GluVal: 3.528 ± 0.027	0.892GluTrp: 0.892 ± 0.013	1.585GluTyr: 1.585 ± 0.017	0.0GluXaa: 0.0 ± 0.0
Phe	3.229PheAla: 3.229 ± 0.027	0.649PheCys: 0.649 ± 0.011	2.263PheAsp: 2.263 ± 0.02	2.185PheGlu: 2.185 ± 0.021	1.603PhePhe: 1.603 ± 0.021	2.89PheGly: 2.89 ± 0.033	0.965PheHis: 0.965 ± 0.013	1.662PheIle: 1.662 ± 0.021	1.493PheLys: 1.493 ± 0.017	3.36PheLeu: 3.36 ± 0.028	0.782PheMet: 0.782 ± 0.012	1.354PheAsn: 1.354 ± 0.016	1.866PhePro: 1.866 ± 0.019	1.376PheGln: 1.376 ± 0.016	1.992PheArg: 1.992 ± 0.02	2.844PheSer: 2.844 ± 0.024	2.09PheThr: 2.09 ± 0.02	2.299PheVal: 2.299 ± 0.024	0.666PheTrp: 0.666 ± 0.014	1.079PheTyr: 1.079 ± 0.015	0.0PheXaa: 0.0 ± 0.0
Gly	5.267GlyAla: 5.267 ± 0.039	0.971GlyCys: 0.971 ± 0.016	3.502GlyAsp: 3.502 ± 0.025	3.7GlyGlu: 3.7 ± 0.028	2.689GlyPhe: 2.689 ± 0.024	5.607GlyGly: 5.607 ± 0.055	1.726GlyHis: 1.726 ± 0.021	3.478GlyIle: 3.478 ± 0.03	3.595GlyLys: 3.595 ± 0.029	5.847GlyLeu: 5.847 ± 0.037	1.739GlyMet: 1.739 ± 0.02	2.476GlyAsn: 2.476 ± 0.024	3.066GlyPro: 3.066 ± 0.027	2.522GlyGln: 2.522 ± 0.022	4.334GlyArg: 4.334 ± 0.032	5.433GlySer: 5.433 ± 0.039	3.847GlyThr: 3.847 ± 0.028	4.17GlyVal: 4.17 ± 0.03	1.195GlyTrp: 1.195 ± 0.015	2.057GlyTyr: 2.057 ± 0.023	0.0GlyXaa: 0.0 ± 0.0
His	2.239HisAla: 2.239 ± 0.019	0.454HisCys: 0.454 ± 0.01	1.464HisAsp: 1.464 ± 0.017	1.487HisGlu: 1.487 ± 0.017	1.045HisPhe: 1.045 ± 0.015	1.799HisGly: 1.799 ± 0.02	0.999HisHis: 0.999 ± 0.016	1.282HisIle: 1.282 ± 0.016	1.004HisLys: 1.004 ± 0.013	2.421HisLeu: 2.421 ± 0.022	0.541HisMet: 0.541 ± 0.01	0.879HisAsn: 0.879 ± 0.012	1.706HisPro: 1.706 ± 0.017	1.078HisGln: 1.078 ± 0.015	1.718HisArg: 1.718 ± 0.018	2.02HisSer: 2.02 ± 0.021	1.452HisThr: 1.452 ± 0.019	1.581HisVal: 1.581 ± 0.016	0.398HisTrp: 0.398 ± 0.009	0.73HisTyr: 0.73 ± 0.012	0.0HisXaa: 0.0 ± 0.0
Ile	4.396IleAla: 4.396 ± 0.035	0.844IleCys: 0.844 ± 0.013	2.86IleAsp: 2.86 ± 0.021	2.835IleGlu: 2.835 ± 0.022	1.936IlePhe: 1.936 ± 0.02	3.173IleGly: 3.173 ± 0.027	1.267IleHis: 1.267 ± 0.017	2.36IleIle: 2.36 ± 0.023	2.15IleLys: 2.15 ± 0.021	4.417IleLeu: 4.417 ± 0.034	1.047IleMet: 1.047 ± 0.015	1.75IleAsn: 1.75 ± 0.019	2.914IlePro: 2.914 ± 0.028	1.814IleGln: 1.814 ± 0.017	2.764IleArg: 2.764 ± 0.022	3.809IleSer: 3.809 ± 0.029	2.805IleThr: 2.805 ± 0.021	3.029IleVal: 3.029 ± 0.025	0.753IleTrp: 0.753 ± 0.013	1.374IleTyr: 1.374 ± 0.017	0.0IleXaa: 0.0 ± 0.0
Lys	4.651LysAla: 4.651 ± 0.04	0.562LysCys: 0.562 ± 0.012	2.84LysAsp: 2.84 ± 0.028	3.361LysGlu: 3.361 ± 0.029	1.454LysPhe: 1.454 ± 0.017	2.917LysGly: 2.917 ± 0.026	1.314LysHis: 1.314 ± 0.013	2.304LysIle: 2.304 ± 0.022	3.44LysLys: 3.44 ± 0.039	4.161LysLeu: 4.161 ± 0.032	1.08LysMet: 1.08 ± 0.016	1.768LysAsn: 1.768 ± 0.016	2.703LysPro: 2.703 ± 0.024	2.139LysGln: 2.139 ± 0.02	3.602LysArg: 3.602 ± 0.03	3.571LysSer: 3.571 ± 0.025	2.856LysThr: 2.856 ± 0.022	2.794LysVal: 2.794 ± 0.027	0.728LysTrp: 0.728 ± 0.011	1.346LysTyr: 1.346 ± 0.017	0.0LysXaa: 0.0 ± 0.0
Leu	8.011LeuAla: 8.011 ± 0.043	1.375LeuCys: 1.375 ± 0.016	4.922LeuAsp: 4.922 ± 0.033	5.397LeuGlu: 5.397 ± 0.042	3.146LeuPhe: 3.146 ± 0.032	5.704LeuGly: 5.704 ± 0.033	2.521LeuHis: 2.521 ± 0.022	3.897LeuIle: 3.897 ± 0.032	4.236LeuLys: 4.236 ± 0.033	8.307LeuLeu: 8.307 ± 0.06	1.789LeuMet: 1.789 ± 0.019	3.065LeuAsn: 3.065 ± 0.024	5.58LeuPro: 5.58 ± 0.035	3.997LeuGln: 3.997 ± 0.034	6.037LeuArg: 6.037 ± 0.042	6.887LeuSer: 6.887 ± 0.044	4.847LeuThr: 4.847 ± 0.032	5.139LeuVal: 5.139 ± 0.038	1.221LeuTrp: 1.221 ± 0.015	2.317LeuTyr: 2.317 ± 0.023	0.0LeuXaa: 0.0 ± 0.0
Met	2.232MetAla: 2.232 ± 0.02	0.291MetCys: 0.291 ± 0.008	1.194MetAsp: 1.194 ± 0.012	1.273MetGlu: 1.273 ± 0.017	0.739MetPhe: 0.739 ± 0.012	1.38MetGly: 1.38 ± 0.017	0.596MetHis: 0.596 ± 0.01	1.04MetIle: 1.04 ± 0.014	1.057MetLys: 1.057 ± 0.016	2.068MetLeu: 2.068 ± 0.02	0.59MetMet: 0.59 ± 0.011	0.837MetAsn: 0.837 ± 0.014	1.408MetPro: 1.408 ± 0.018	0.991MetGln: 0.991 ± 0.014	1.462MetArg: 1.462 ± 0.018	1.848MetSer: 1.848 ± 0.018	1.301MetThr: 1.301 ± 0.017	1.226MetVal: 1.226 ± 0.015	0.269MetTrp: 0.269 ± 0.006	0.561MetTyr: 0.561 ± 0.009	0.0MetXaa: 0.0 ± 0.0
Asn	3.481AsnAla: 3.481 ± 0.025	0.478AsnCys: 0.478 ± 0.01	1.986AsnAsp: 1.986 ± 0.02	2.014AsnGlu: 2.014 ± 0.021	1.436AsnPhe: 1.436 ± 0.019	3.052AsnGly: 3.052 ± 0.033	0.937AsnHis: 0.937 ± 0.012	1.913AsnIle: 1.913 ± 0.021	1.481AsnLys: 1.481 ± 0.016	3.131AsnLeu: 3.131 ± 0.026	0.8AsnMet: 0.8 ± 0.012	1.388AsnAsn: 1.388 ± 0.021	2.199AsnPro: 2.199 ± 0.024	1.33AsnGln: 1.33 ± 0.017	1.902AsnArg: 1.902 ± 0.02	2.627AsnSer: 2.627 ± 0.023	2.193AsnThr: 2.193 ± 0.023	2.296AsnVal: 2.296 ± 0.026	0.552AsnTrp: 0.552 ± 0.01	0.979AsnTyr: 0.979 ± 0.013	0.0AsnXaa: 0.0 ± 0.0
Pro	5.267ProAla: 5.267 ± 0.04	0.646ProCys: 0.646 ± 0.011	3.011ProAsp: 3.011 ± 0.024	3.623ProGlu: 3.623 ± 0.029	1.987ProPhe: 1.987 ± 0.02	3.728ProGly: 3.728 ± 0.028	1.452ProHis: 1.452 ± 0.016	2.451ProIle: 2.451 ± 0.021	2.765ProLys: 2.765 ± 0.026	4.492ProLeu: 4.492 ± 0.035	1.095ProMet: 1.095 ± 0.014	2.192ProAsn: 2.192 ± 0.023	4.959ProPro: 4.959 ± 0.067	2.444ProGln: 2.444 ± 0.024	3.424ProArg: 3.424 ± 0.03	5.593ProSer: 5.593 ± 0.049	3.77ProThr: 3.77 ± 0.033	3.181ProVal: 3.181 ± 0.024	0.749ProTrp: 0.749 ± 0.012	1.524ProTyr: 1.524 ± 0.018	0.0ProXaa: 0.0 ± 0.0
Gln	3.79GlnAla: 3.79 ± 0.034	0.535GlnCys: 0.535 ± 0.011	2.177GlnAsp: 2.177 ± 0.02	2.458GlnGlu: 2.458 ± 0.024	1.204GlnPhe: 1.204 ± 0.015	2.352GlnGly: 2.352 ± 0.02	1.303GlnHis: 1.303 ± 0.014	1.934GlnIle: 1.934 ± 0.02	2.145GlnLys: 2.145 ± 0.019	3.531GlnLeu: 3.531 ± 0.028	0.941GlnMet: 0.941 ± 0.016	1.685GlnAsn: 1.685 ± 0.019	2.574GlnPro: 2.574 ± 0.03	2.597GlnGln: 2.597 ± 0.041	2.914GlnArg: 2.914 ± 0.024	3.119GlnSer: 3.119 ± 0.027	2.404GlnThr: 2.404 ± 0.024	2.185GlnVal: 2.185 ± 0.018	0.595GlnTrp: 0.595 ± 0.012	1.258GlnTyr: 1.258 ± 0.015	0.0GlnXaa: 0.0 ± 0.0
Arg	5.052ArgAla: 5.052 ± 0.035	0.921ArgCys: 0.921 ± 0.015	3.425ArgAsp: 3.425 ± 0.028	3.926ArgGlu: 3.926 ± 0.033	2.192ArgPhe: 2.192 ± 0.02	3.713ArgGly: 3.713 ± 0.033	1.8ArgHis: 1.8 ± 0.018	3.041ArgIle: 3.041 ± 0.028	3.798ArgLys: 3.798 ± 0.03	5.683ArgLeu: 5.683 ± 0.039	1.497ArgMet: 1.497 ± 0.017	2.389ArgAsn: 2.389 ± 0.02	3.628ArgPro: 3.628 ± 0.031	2.87ArgGln: 2.87 ± 0.027	5.413ArgArg: 5.413 ± 0.044	5.068ArgSer: 5.068 ± 0.04	3.527ArgThr: 3.527 ± 0.031	3.337ArgVal: 3.337 ± 0.025	1.018ArgTrp: 1.018 ± 0.014	1.71ArgTyr: 1.71 ± 0.018	0.0ArgXaa: 0.0 ± 0.0
Ser	6.856SerAla: 6.856 ± 0.044	1.081SerCys: 1.081 ± 0.016	4.041SerAsp: 4.041 ± 0.035	4.059SerGlu: 4.059 ± 0.031	2.853SerPhe: 2.853 ± 0.022	5.421SerGly: 5.421 ± 0.035	2.044SerHis: 2.044 ± 0.022	3.967SerIle: 3.967 ± 0.026	3.821SerLys: 3.821 ± 0.027	6.858SerLeu: 6.858 ± 0.037	1.799SerMet: 1.799 ± 0.02	2.983SerAsn: 2.983 ± 0.025	5.066SerPro: 5.066 ± 0.051	3.212SerGln: 3.212 ± 0.024	5.258SerArg: 5.258 ± 0.043	8.159SerSer: 8.159 ± 0.066	5.457SerThr: 5.457 ± 0.038	4.23SerVal: 4.23 ± 0.032	1.196SerTrp: 1.196 ± 0.015	2.045SerTyr: 2.045 ± 0.022	0.0SerXaa: 0.0 ± 0.0
Thr	5.444ThrAla: 5.444 ± 0.033	0.888ThrCys: 0.888 ± 0.015	2.675ThrAsp: 2.675 ± 0.023	2.863ThrGlu: 2.863 ± 0.023	2.313ThrPhe: 2.313 ± 0.022	3.922ThrGly: 3.922 ± 0.029	1.395ThrHis: 1.395 ± 0.017	3.124ThrIle: 3.124 ± 0.021	2.643ThrLys: 2.643 ± 0.025	5.225ThrLeu: 5.225 ± 0.038	1.22ThrMet: 1.22 ± 0.015	2.112ThrAsn: 2.112 ± 0.021	4.15ThrPro: 4.15 ± 0.035	2.113ThrGln: 2.113 ± 0.022	3.308ThrArg: 3.308 ± 0.023	5.395ThrSer: 5.395 ± 0.04	4.23ThrThr: 4.23 ± 0.037	3.396ThrVal: 3.396 ± 0.029	0.878ThrTrp: 0.878 ± 0.014	1.608ThrTyr: 1.608 ± 0.019	0.0ThrXaa: 0.0 ± 0.0
Val	5.122ValAla: 5.122 ± 0.033	0.914ValCys: 0.914 ± 0.012	3.444ValAsp: 3.444 ± 0.029	3.717ValGlu: 3.717 ± 0.028	2.274ValPhe: 2.274 ± 0.021	3.889ValGly: 3.889 ± 0.033	1.476ValHis: 1.476 ± 0.017	2.72ValIle: 2.72 ± 0.028	2.937ValLys: 2.937 ± 0.022	5.433ValLeu: 5.433 ± 0.032	1.258ValMet: 1.258 ± 0.014	2.133ValAsn: 2.133 ± 0.021	3.286ValPro: 3.286 ± 0.025	2.426ValGln: 2.426 ± 0.024	3.527ValArg: 3.527 ± 0.027	4.314ValSer: 4.314 ± 0.028	3.267ValThr: 3.267 ± 0.029	4.085ValVal: 4.085 ± 0.036	0.908ValTrp: 0.908 ± 0.014	1.634ValTyr: 1.634 ± 0.021	0.0ValXaa: 0.0 ± 0.0
Trp	1.113TrpAla: 1.113 ± 0.014	0.237TrpCys: 0.237 ± 0.007	0.845TrpAsp: 0.845 ± 0.013	0.81TrpGlu: 0.81 ± 0.013	0.546TrpPhe: 0.546 ± 0.01	0.832TrpGly: 0.832 ± 0.016	0.421TrpHis: 0.421 ± 0.01	0.815TrpIle: 0.815 ± 0.014	0.845TrpLys: 0.845 ± 0.012	1.453TrpLeu: 1.453 ± 0.019	0.391TrpMet: 0.391 ± 0.009	0.656TrpAsn: 0.656 ± 0.01	0.71TrpPro: 0.71 ± 0.011	0.699TrpGln: 0.699 ± 0.01	1.119TrpArg: 1.119 ± 0.013	1.169TrpSer: 1.169 ± 0.016	0.996TrpThr: 0.996 ± 0.014	0.811TrpVal: 0.811 ± 0.012	0.282TrpTrp: 0.282 ± 0.008	0.474TrpTyr: 0.474 ± 0.009	0.0TrpXaa: 0.0 ± 0.0
Tyr	2.269TyrAla: 2.269 ± 0.023	0.468TyrCys: 0.468 ± 0.01	1.64TyrAsp: 1.64 ± 0.017	1.549TyrGlu: 1.549 ± 0.015	1.15TyrPhe: 1.15 ± 0.014	2.114TyrGly: 2.114 ± 0.025	0.791TyrHis: 0.791 ± 0.012	1.381TyrIle: 1.381 ± 0.016	1.094TyrLys: 1.094 ± 0.016	2.519TyrLeu: 2.519 ± 0.027	0.605TyrMet: 0.605 ± 0.011	1.102TyrAsn: 1.102 ± 0.015	1.416TyrPro: 1.416 ± 0.021	1.141TyrGln: 1.141 ± 0.014	1.618TyrArg: 1.618 ± 0.017	2.009TyrSer: 2.009 ± 0.023	1.621TyrThr: 1.621 ± 0.019	1.6TyrVal: 1.6 ± 0.018	0.443TyrTrp: 0.443 ± 0.01	0.907TyrTyr: 0.907 ± 0.016	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.0XaaAla: 0.0 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.0XaaAsp: 0.0 ± 0.0	0.0XaaGlu: 0.0 ± 0.0	0.0XaaPhe: 0.0 ± 0.0	0.0XaaGly: 0.0 ± 0.0	0.0XaaHis: 0.0 ± 0.0	0.0XaaIle: 0.0 ± 0.0	0.0XaaLys: 0.0 ± 0.0	0.0XaaLeu: 0.0 ± 0.0	0.0XaaMet: 0.0 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.0XaaPro: 0.0 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.0XaaArg: 0.0 ± 0.0	0.0XaaSer: 0.0 ± 0.0	0.0XaaThr: 0.0 ± 0.0	0.0XaaVal: 0.0 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.0XaaXaa: 0.0 ± 0.0

Statistics based on 13062 proteins (5586011 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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