Amino acid dipepetide frequency for Oryza meridionalis

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	15.036AlaAla: 15.036 ± 0.075	1.556AlaCys: 1.556 ± 0.011	4.272AlaAsp: 4.272 ± 0.019	4.901AlaGlu: 4.901 ± 0.023	2.887AlaPhe: 2.887 ± 0.018	6.714AlaGly: 6.714 ± 0.026	1.73AlaHis: 1.73 ± 0.012	3.672AlaIle: 3.672 ± 0.015	3.695AlaLys: 3.695 ± 0.022	7.648AlaLeu: 7.648 ± 0.024	2.317AlaMet: 2.317 ± 0.013	2.614AlaAsn: 2.614 ± 0.013	4.746AlaPro: 4.746 ± 0.025	2.39AlaGln: 2.39 ± 0.011	5.512AlaArg: 5.512 ± 0.025	7.528AlaSer: 7.528 ± 0.029	4.844AlaThr: 4.844 ± 0.018	6.794AlaVal: 6.794 ± 0.024	1.013AlaTrp: 1.013 ± 0.008	1.871AlaTyr: 1.871 ± 0.013	0.003AlaXaa: 0.003 ± 0.0
Cys	1.343CysAla: 1.343 ± 0.01	0.605CysCys: 0.605 ± 0.008	0.902CysAsp: 0.902 ± 0.008	0.831CysGlu: 0.831 ± 0.009	0.816CysPhe: 0.816 ± 0.007	1.523CysGly: 1.523 ± 0.012	0.529CysHis: 0.529 ± 0.006	0.914CysIle: 0.914 ± 0.008	0.925CysLys: 0.925 ± 0.009	1.792CysLeu: 1.792 ± 0.013	0.448CysMet: 0.448 ± 0.006	0.75CysAsn: 0.75 ± 0.008	1.016CysPro: 1.016 ± 0.01	0.584CysGln: 0.584 ± 0.007	1.288CysArg: 1.288 ± 0.01	1.889CysSer: 1.889 ± 0.011	0.921CysThr: 0.921 ± 0.008	1.122CysVal: 1.122 ± 0.008	0.266CysTrp: 0.266 ± 0.004	0.519CysTyr: 0.519 ± 0.006	0.0CysXaa: 0.0 ± 0.0
Asp	4.703AspAla: 4.703 ± 0.022	0.901AspCys: 0.901 ± 0.008	4.399AspAsp: 4.399 ± 0.024	3.949AspGlu: 3.949 ± 0.016	1.965AspPhe: 1.965 ± 0.011	4.767AspGly: 4.767 ± 0.022	1.305AspHis: 1.305 ± 0.009	2.682AspIle: 2.682 ± 0.017	2.404AspLys: 2.404 ± 0.015	4.83AspLeu: 4.83 ± 0.02	1.411AspMet: 1.411 ± 0.01	1.834AspAsn: 1.834 ± 0.012	2.657AspPro: 2.657 ± 0.014	1.597AspGln: 1.597 ± 0.011	2.685AspArg: 2.685 ± 0.013	3.746AspSer: 3.746 ± 0.017	2.215AspThr: 2.215 ± 0.012	3.766AspVal: 3.766 ± 0.016	0.683AspTrp: 0.683 ± 0.007	1.394AspTyr: 1.394 ± 0.01	0.001AspXaa: 0.001 ± 0.0
Glu	5.353GluAla: 5.353 ± 0.02	0.862GluCys: 0.862 ± 0.009	3.618GluAsp: 3.618 ± 0.017	5.927GluGlu: 5.927 ± 0.032	1.987GluPhe: 1.987 ± 0.011	3.734GluGly: 3.734 ± 0.019	1.41GluHis: 1.41 ± 0.01	2.995GluIle: 2.995 ± 0.017	3.711GluLys: 3.711 ± 0.021	5.859GluLeu: 5.859 ± 0.024	1.653GluMet: 1.653 ± 0.013	2.287GluAsn: 2.287 ± 0.013	2.285GluPro: 2.285 ± 0.015	2.209GluGln: 2.209 ± 0.012	3.671GluArg: 3.671 ± 0.018	3.792GluSer: 3.792 ± 0.018	2.586GluThr: 2.586 ± 0.014	4.061GluVal: 4.061 ± 0.018	0.758GluTrp: 0.758 ± 0.007	1.49GluTyr: 1.49 ± 0.01	0.001GluXaa: 0.001 ± 0.0
Phe	2.783PheAla: 2.783 ± 0.014	0.769PheCys: 0.769 ± 0.007	2.113PheAsp: 2.113 ± 0.011	1.822PheGlu: 1.822 ± 0.012	1.671PhePhe: 1.671 ± 0.013	2.849PheGly: 2.849 ± 0.019	0.982PheHis: 0.982 ± 0.008	1.611PheIle: 1.611 ± 0.01	1.48PheLys: 1.48 ± 0.012	3.791PheLeu: 3.791 ± 0.016	0.863PheMet: 0.863 ± 0.007	1.287PheAsn: 1.287 ± 0.01	1.903PhePro: 1.903 ± 0.012	1.226PheGln: 1.226 ± 0.01	2.01PheArg: 2.01 ± 0.011	3.309PheSer: 3.309 ± 0.015	1.744PheThr: 1.744 ± 0.011	2.549PheVal: 2.549 ± 0.013	0.509PheTrp: 0.509 ± 0.006	1.043PheTyr: 1.043 ± 0.007	0.001PheXaa: 0.001 ± 0.0
Gly	6.174GlyAla: 6.174 ± 0.028	1.498GlyCys: 1.498 ± 0.013	4.286GlyAsp: 4.286 ± 0.019	4.329GlyGlu: 4.329 ± 0.019	2.746GlyPhe: 2.746 ± 0.015	9.427GlyGly: 9.427 ± 0.058	1.769GlyHis: 1.769 ± 0.012	3.132GlyIle: 3.132 ± 0.017	3.564GlyLys: 3.564 ± 0.016	5.775GlyLeu: 5.775 ± 0.023	1.665GlyMet: 1.665 ± 0.011	2.737GlyAsn: 2.737 ± 0.015	2.68GlyPro: 2.68 ± 0.014	2.157GlyGln: 2.157 ± 0.012	4.941GlyArg: 4.941 ± 0.021	6.15GlySer: 6.15 ± 0.025	3.368GlyThr: 3.368 ± 0.017	4.768GlyVal: 4.768 ± 0.02	1.038GlyTrp: 1.038 ± 0.009	1.996GlyTyr: 1.996 ± 0.013	0.002GlyXaa: 0.002 ± 0.0
His	1.933HisAla: 1.933 ± 0.011	0.539HisCys: 0.539 ± 0.005	1.261HisAsp: 1.261 ± 0.01	1.246HisGlu: 1.246 ± 0.01	0.929HisPhe: 0.929 ± 0.008	2.153HisGly: 2.153 ± 0.013	1.166HisHis: 1.166 ± 0.013	1.095HisIle: 1.095 ± 0.009	0.979HisLys: 0.979 ± 0.008	2.578HisLeu: 2.578 ± 0.016	0.566HisMet: 0.566 ± 0.006	0.824HisAsn: 0.824 ± 0.007	1.553HisPro: 1.553 ± 0.011	0.97HisGln: 0.97 ± 0.009	1.812HisArg: 1.812 ± 0.011	1.729HisSer: 1.729 ± 0.012	0.986HisThr: 0.986 ± 0.008	1.624HisVal: 1.624 ± 0.009	0.294HisTrp: 0.294 ± 0.005	0.657HisTyr: 0.657 ± 0.007	0.001HisXaa: 0.001 ± 0.0
Ile	3.54IleAla: 3.54 ± 0.016	0.98IleCys: 0.98 ± 0.008	2.517IleAsp: 2.517 ± 0.014	2.481IleGlu: 2.481 ± 0.014	1.766IlePhe: 1.766 ± 0.012	3.041IleGly: 3.041 ± 0.017	1.153IleHis: 1.153 ± 0.01	2.233IleIle: 2.233 ± 0.015	2.268IleLys: 2.268 ± 0.014	4.339IleLeu: 4.339 ± 0.02	0.981IleMet: 0.981 ± 0.009	1.694IleAsn: 1.694 ± 0.01	2.557IlePro: 2.557 ± 0.017	1.575IleGln: 1.575 ± 0.011	2.413IleArg: 2.413 ± 0.013	3.949IleSer: 3.949 ± 0.018	2.27IleThr: 2.27 ± 0.012	2.994IleVal: 2.994 ± 0.015	0.609IleTrp: 0.609 ± 0.007	1.248IleTyr: 1.248 ± 0.009	0.001IleXaa: 0.001 ± 0.0
Lys	3.705LysAla: 3.705 ± 0.019	0.803LysCys: 0.803 ± 0.008	2.653LysAsp: 2.653 ± 0.013	3.659LysGlu: 3.659 ± 0.023	1.629LysPhe: 1.629 ± 0.01	2.998LysGly: 2.998 ± 0.016	1.155LysHis: 1.155 ± 0.009	2.53LysIle: 2.53 ± 0.014	3.615LysLys: 3.615 ± 0.026	4.854LysLeu: 4.854 ± 0.023	1.231LysMet: 1.231 ± 0.009	1.984LysAsn: 1.984 ± 0.013	2.306LysPro: 2.306 ± 0.015	1.883LysGln: 1.883 ± 0.012	3.133LysArg: 3.133 ± 0.015	3.438LysSer: 3.438 ± 0.018	2.209LysThr: 2.209 ± 0.014	3.137LysVal: 3.137 ± 0.015	0.587LysTrp: 0.587 ± 0.007	1.313LysTyr: 1.313 ± 0.01	0.001LysXaa: 0.001 ± 0.0
Leu	8.096LeuAla: 8.096 ± 0.028	1.783LeuCys: 1.783 ± 0.012	5.011LeuAsp: 5.011 ± 0.019	5.629LeuGlu: 5.629 ± 0.025	3.372LeuPhe: 3.372 ± 0.017	5.856LeuGly: 5.856 ± 0.02	2.774LeuHis: 2.774 ± 0.015	3.688LeuIle: 3.688 ± 0.018	4.663LeuLys: 4.663 ± 0.02	10.451LeuLeu: 10.451 ± 0.038	2.001LeuMet: 2.001 ± 0.012	3.075LeuAsn: 3.075 ± 0.016	5.955LeuPro: 5.955 ± 0.025	4.03LeuGln: 4.03 ± 0.019	6.351LeuArg: 6.351 ± 0.023	7.844LeuSer: 7.844 ± 0.029	4.209LeuThr: 4.209 ± 0.019	6.368LeuVal: 6.368 ± 0.022	1.124LeuTrp: 1.124 ± 0.008	2.276LeuTyr: 2.276 ± 0.012	0.002LeuXaa: 0.002 ± 0.0
Met	2.771MetAla: 2.771 ± 0.014	0.344MetCys: 0.344 ± 0.005	1.455MetAsp: 1.455 ± 0.009	1.865MetGlu: 1.865 ± 0.011	0.761MetPhe: 0.761 ± 0.007	1.536MetGly: 1.536 ± 0.01	0.565MetHis: 0.565 ± 0.006	0.986MetIle: 0.986 ± 0.009	1.262MetLys: 1.262 ± 0.011	2.216MetLeu: 2.216 ± 0.014	0.673MetMet: 0.673 ± 0.007	0.855MetAsn: 0.855 ± 0.007	1.282MetPro: 1.282 ± 0.01	0.9MetGln: 0.9 ± 0.008	1.371MetArg: 1.371 ± 0.009	1.767MetSer: 1.767 ± 0.011	1.048MetThr: 1.048 ± 0.008	1.69MetVal: 1.69 ± 0.01	0.291MetTrp: 0.291 ± 0.004	0.587MetTyr: 0.587 ± 0.007	0.0MetXaa: 0.0 ± 0.0
Asn	2.616AsnAla: 2.616 ± 0.014	0.722AsnCys: 0.722 ± 0.007	1.755AsnAsp: 1.755 ± 0.01	1.88AsnGlu: 1.88 ± 0.013	1.412AsnPhe: 1.412 ± 0.011	2.84AsnGly: 2.84 ± 0.015	0.893AsnHis: 0.893 ± 0.008	1.996AsnIle: 1.996 ± 0.011	1.816AsnLys: 1.816 ± 0.012	3.758AsnLeu: 3.758 ± 0.026	0.989AsnMet: 0.989 ± 0.009	1.732AsnAsn: 1.732 ± 0.015	1.891AsnPro: 1.891 ± 0.012	1.294AsnGln: 1.294 ± 0.011	1.752AsnArg: 1.752 ± 0.01	2.907AsnSer: 2.907 ± 0.017	1.702AsnThr: 1.702 ± 0.011	2.228AsnVal: 2.228 ± 0.013	0.452AsnTrp: 0.452 ± 0.006	0.999AsnTyr: 0.999 ± 0.008	0.001AsnXaa: 0.001 ± 0.0
Pro	5.312ProAla: 5.312 ± 0.024	0.966ProCys: 0.966 ± 0.01	2.678ProAsp: 2.678 ± 0.014	3.183ProGlu: 3.183 ± 0.016	1.891ProPhe: 1.891 ± 0.012	3.293ProGly: 3.293 ± 0.017	1.275ProHis: 1.275 ± 0.009	1.969ProIle: 1.969 ± 0.012	2.256ProLys: 2.256 ± 0.015	4.693ProLeu: 4.693 ± 0.021	1.107ProMet: 1.107 ± 0.009	1.849ProAsn: 1.849 ± 0.014	6.21ProPro: 6.21 ± 0.043	1.781ProGln: 1.781 ± 0.013	3.531ProArg: 3.531 ± 0.022	5.615ProSer: 5.615 ± 0.026	2.865ProThr: 2.865 ± 0.014	3.428ProVal: 3.428 ± 0.018	0.706ProTrp: 0.706 ± 0.007	1.271ProTyr: 1.271 ± 0.01	0.002ProXaa: 0.002 ± 0.0
Gln	2.59GlnAla: 2.59 ± 0.013	0.592GlnCys: 0.592 ± 0.006	1.589GlnAsp: 1.589 ± 0.011	2.242GlnGlu: 2.242 ± 0.014	1.172GlnPhe: 1.172 ± 0.009	2.111GlnGly: 2.111 ± 0.012	0.977GlnHis: 0.977 ± 0.009	1.641GlnIle: 1.641 ± 0.01	1.854GlnLys: 1.854 ± 0.014	3.505GlnLeu: 3.505 ± 0.016	0.926GlnMet: 0.926 ± 0.008	1.365GlnAsn: 1.365 ± 0.011	1.942GlnPro: 1.942 ± 0.014	2.29GlnGln: 2.29 ± 0.027	2.238GlnArg: 2.238 ± 0.012	2.451GlnSer: 2.451 ± 0.013	1.424GlnThr: 1.424 ± 0.011	2.111GlnVal: 2.111 ± 0.012	0.47GlnTrp: 0.47 ± 0.005	0.88GlnTyr: 0.88 ± 0.008	0.001GlnXaa: 0.001 ± 0.0
Arg	5.09ArgAla: 5.09 ± 0.025	1.367ArgCys: 1.367 ± 0.01	2.939ArgAsp: 2.939 ± 0.015	3.533ArgGlu: 3.533 ± 0.017	2.204ArgPhe: 2.204 ± 0.015	4.49ArgGly: 4.49 ± 0.021	1.723ArgHis: 1.723 ± 0.011	2.638ArgIle: 2.638 ± 0.013	3.31ArgLys: 3.31 ± 0.017	6.046ArgLeu: 6.046 ± 0.024	1.522ArgMet: 1.522 ± 0.01	2.106ArgAsn: 2.106 ± 0.013	3.389ArgPro: 3.389 ± 0.019	2.1ArgGln: 2.1 ± 0.011	7.319ArgArg: 7.319 ± 0.039	5.021ArgSer: 5.021 ± 0.02	2.67ArgThr: 2.67 ± 0.013	3.748ArgVal: 3.748 ± 0.016	1.088ArgTrp: 1.088 ± 0.009	1.571ArgTyr: 1.571 ± 0.012	0.002ArgXaa: 0.002 ± 0.0
Ser	6.586SerAla: 6.586 ± 0.022	1.747SerCys: 1.747 ± 0.012	4.093SerAsp: 4.093 ± 0.018	4.035SerGlu: 4.035 ± 0.018	3.305SerPhe: 3.305 ± 0.015	6.088SerGly: 6.088 ± 0.026	1.85SerHis: 1.85 ± 0.012	3.713SerIle: 3.713 ± 0.017	3.791SerLys: 3.791 ± 0.019	7.842SerLeu: 7.842 ± 0.032	1.996SerMet: 1.996 ± 0.012	3.158SerAsn: 3.158 ± 0.017	5.278SerPro: 5.278 ± 0.028	2.496SerGln: 2.496 ± 0.014	4.865SerArg: 4.865 ± 0.021	10.716SerSer: 10.716 ± 0.045	4.574SerThr: 4.574 ± 0.018	4.86SerVal: 4.86 ± 0.02	1.17SerTrp: 1.17 ± 0.01	2.112SerTyr: 2.112 ± 0.013	0.002SerXaa: 0.002 ± 0.0
Thr	4.489ThrAla: 4.489 ± 0.018	0.922ThrCys: 0.922 ± 0.008	2.293ThrAsp: 2.293 ± 0.012	2.581ThrGlu: 2.581 ± 0.016	1.727ThrPhe: 1.727 ± 0.01	3.495ThrGly: 3.495 ± 0.017	0.978ThrHis: 0.978 ± 0.008	2.333ThrIle: 2.333 ± 0.014	2.217ThrLys: 2.217 ± 0.013	4.191ThrLeu: 4.191 ± 0.017	1.259ThrMet: 1.259 ± 0.01	1.772ThrAsn: 1.772 ± 0.012	2.862ThrPro: 2.862 ± 0.016	1.345ThrGln: 1.345 ± 0.009	2.627ThrArg: 2.627 ± 0.014	4.369ThrSer: 4.369 ± 0.02	3.247ThrThr: 3.247 ± 0.019	3.452ThrVal: 3.452 ± 0.017	0.612ThrTrp: 0.612 ± 0.008	1.218ThrTyr: 1.218 ± 0.01	0.001ThrXaa: 0.001 ± 0.0
Val	6.778ValAla: 6.778 ± 0.027	1.168ValCys: 1.168 ± 0.01	3.934ValAsp: 3.934 ± 0.016	4.103ValGlu: 4.103 ± 0.017	2.485ValPhe: 2.485 ± 0.015	4.508ValGly: 4.508 ± 0.021	1.642ValHis: 1.642 ± 0.011	2.894ValIle: 2.894 ± 0.014	3.077ValLys: 3.077 ± 0.017	6.519ValLeu: 6.519 ± 0.023	1.534ValMet: 1.534 ± 0.01	2.078ValAsn: 2.078 ± 0.013	3.554ValPro: 3.554 ± 0.013	2.249ValGln: 2.249 ± 0.01	3.72ValArg: 3.72 ± 0.017	5.062ValSer: 5.062 ± 0.018	3.262ValThr: 3.262 ± 0.015	5.696ValVal: 5.696 ± 0.022	0.792ValTrp: 0.792 ± 0.008	1.76ValTyr: 1.76 ± 0.013	0.001ValXaa: 0.001 ± 0.0
Trp	0.977TrpAla: 0.977 ± 0.009	0.265TrpCys: 0.265 ± 0.004	0.679TrpAsp: 0.679 ± 0.007	0.726TrpGlu: 0.726 ± 0.007	0.488TrpPhe: 0.488 ± 0.006	0.778TrpGly: 0.778 ± 0.008	0.317TrpHis: 0.317 ± 0.004	0.598TrpIle: 0.598 ± 0.006	0.75TrpLys: 0.75 ± 0.007	1.291TrpLeu: 1.291 ± 0.01	0.368TrpMet: 0.368 ± 0.005	0.561TrpAsn: 0.561 ± 0.006	0.617TrpPro: 0.617 ± 0.007	0.456TrpGln: 0.456 ± 0.006	1.168TrpArg: 1.168 ± 0.009	1.029TrpSer: 1.029 ± 0.008	0.655TrpThr: 0.655 ± 0.007	0.797TrpVal: 0.797 ± 0.008	0.294TrpTrp: 0.294 ± 0.005	0.327TrpTyr: 0.327 ± 0.005	0.0TrpXaa: 0.0 ± 0.0
Tyr	1.863TyrAla: 1.863 ± 0.01	0.592TyrCys: 0.592 ± 0.006	1.445TyrAsp: 1.445 ± 0.01	1.324TyrGlu: 1.324 ± 0.008	1.1TyrPhe: 1.1 ± 0.008	2.0TyrGly: 2.0 ± 0.013	0.706TyrHis: 0.706 ± 0.007	1.288TyrIle: 1.288 ± 0.01	1.183TyrLys: 1.183 ± 0.009	2.579TyrLeu: 2.579 ± 0.014	0.713TyrMet: 0.713 ± 0.007	1.072TyrAsn: 1.072 ± 0.009	1.198TyrPro: 1.198 ± 0.009	0.839TyrGln: 0.839 ± 0.008	1.465TyrArg: 1.465 ± 0.011	1.964TyrSer: 1.964 ± 0.011	1.194TyrThr: 1.194 ± 0.01	1.621TyrVal: 1.621 ± 0.011	0.383TyrTrp: 0.383 ± 0.006	0.916TyrTyr: 0.916 ± 0.008	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.003XaaAla: 0.003 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.001XaaAsp: 0.001 ± 0.0	0.001XaaGlu: 0.001 ± 0.0	0.001XaaPhe: 0.001 ± 0.0	0.002XaaGly: 0.002 ± 0.0	0.001XaaHis: 0.001 ± 0.0	0.002XaaIle: 0.002 ± 0.0	0.001XaaLys: 0.001 ± 0.0	0.001XaaLeu: 0.001 ± 0.0	0.001XaaMet: 0.001 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.001XaaPro: 0.001 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.003XaaArg: 0.003 ± 0.0	0.002XaaSer: 0.002 ± 0.0	0.001XaaThr: 0.001 ± 0.0	0.002XaaVal: 0.002 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.053XaaXaa: 0.053 ± 0.021

Statistics based on 40192 proteins (16856824 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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