Amino acid dipepetide frequency for Sus scrofa papillomavirus 2

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
7.106AlaAla: 7.106 ± 2.391
0.395AlaCys: 0.395 ± 0.459
3.553AlaAsp: 3.553 ± 0.468
3.158AlaGlu: 3.158 ± 1.397
0.79AlaPhe: 0.79 ± 0.695
2.369AlaGly: 2.369 ± 0.408
0.79AlaHis: 0.79 ± 0.45
2.369AlaIle: 2.369 ± 0.811
4.343AlaLys: 4.343 ± 1.907
4.343AlaLeu: 4.343 ± 1.339
0.395AlaMet: 0.395 ± 0.625
1.974AlaAsn: 1.974 ± 0.558
4.343AlaPro: 4.343 ± 1.702
2.764AlaGln: 2.764 ± 1.115
4.737AlaArg: 4.737 ± 1.109
5.132AlaSer: 5.132 ± 0.848
5.922AlaThr: 5.922 ± 1.076
2.764AlaVal: 2.764 ± 0.653
0.395AlaTrp: 0.395 ± 0.352
1.579AlaTyr: 1.579 ± 1.068
0.0AlaXaa: 0.0 ± 0.0
Cys
0.79CysAla: 0.79 ± 0.656
1.579CysCys: 1.579 ± 0.655
1.579CysAsp: 1.579 ± 0.545
0.395CysGlu: 0.395 ± 0.516
0.79CysPhe: 0.79 ± 0.427
3.158CysGly: 3.158 ± 1.114
0.79CysHis: 0.79 ± 0.427
1.184CysIle: 1.184 ± 0.61
1.184CysLys: 1.184 ± 0.979
1.579CysLeu: 1.579 ± 1.035
0.0CysMet: 0.0 ± 0.0
0.395CysAsn: 0.395 ± 0.459
2.369CysPro: 2.369 ± 0.596
0.395CysGln: 0.395 ± 0.352
1.579CysArg: 1.579 ± 1.131
2.369CysSer: 2.369 ± 1.222
1.184CysThr: 1.184 ± 0.494
1.579CysVal: 1.579 ± 0.893
0.79CysTrp: 0.79 ± 0.336
0.395CysTyr: 0.395 ± 0.275
0.0CysXaa: 0.0 ± 0.0
Asp
1.184AspAla: 1.184 ± 0.36
1.184AspCys: 1.184 ± 0.316
3.158AspAsp: 3.158 ± 0.941
3.553AspGlu: 3.553 ± 0.559
2.764AspPhe: 2.764 ± 0.782
2.369AspGly: 2.369 ± 0.997
1.184AspHis: 1.184 ± 0.373
3.553AspIle: 3.553 ± 1.08
1.579AspLys: 1.579 ± 0.84
7.896AspLeu: 7.896 ± 1.953
1.579AspMet: 1.579 ± 1.189
2.764AspAsn: 2.764 ± 1.049
7.106AspPro: 7.106 ± 2.379
1.184AspGln: 1.184 ± 0.626
1.579AspArg: 1.579 ± 1.068
7.106AspSer: 7.106 ± 1.762
5.527AspThr: 5.527 ± 1.539
3.948AspVal: 3.948 ± 1.08
0.395AspTrp: 0.395 ± 0.352
1.579AspTyr: 1.579 ± 1.035
0.0AspXaa: 0.0 ± 0.0
Glu
4.737GluAla: 4.737 ± 1.257
0.395GluCys: 0.395 ± 0.275
6.711GluAsp: 6.711 ± 1.9
4.737GluGlu: 4.737 ± 1.197
1.579GluPhe: 1.579 ± 0.495
2.764GluGly: 2.764 ± 1.569
0.395GluHis: 0.395 ± 0.32
1.974GluIle: 1.974 ± 0.551
0.79GluLys: 0.79 ± 0.427
3.948GluLeu: 3.948 ± 0.489
1.974GluMet: 1.974 ± 1.148
3.948GluAsn: 3.948 ± 1.167
3.553GluPro: 3.553 ± 1.49
1.974GluGln: 1.974 ± 1.071
2.764GluArg: 2.764 ± 1.232
5.527GluSer: 5.527 ± 1.15
3.948GluThr: 3.948 ± 1.226
3.553GluVal: 3.553 ± 1.898
0.395GluTrp: 0.395 ± 0.352
1.579GluTyr: 1.579 ± 0.522
0.0GluXaa: 0.0 ± 0.0
Phe
1.579PheAla: 1.579 ± 0.139
0.395PheCys: 0.395 ± 0.459
2.369PheAsp: 2.369 ± 0.848
2.764PheGlu: 2.764 ± 1.301
1.974PhePhe: 1.974 ± 0.928
3.158PheGly: 3.158 ± 1.196
0.395PheHis: 0.395 ± 0.352
1.579PheIle: 1.579 ± 0.672
3.948PheLys: 3.948 ± 2.317
6.317PheLeu: 6.317 ± 0.676
0.79PheMet: 0.79 ± 0.62
1.579PheAsn: 1.579 ± 0.495
1.974PhePro: 1.974 ± 0.638
1.579PheGln: 1.579 ± 1.068
1.184PheArg: 1.184 ± 0.567
1.184PheSer: 1.184 ± 0.826
2.369PheThr: 2.369 ± 0.724
1.184PheVal: 1.184 ± 0.316
0.79PheTrp: 0.79 ± 0.336
1.579PheTyr: 1.579 ± 0.619
0.0PheXaa: 0.0 ± 0.0
Gly
3.948GlyAla: 3.948 ± 1.294
1.974GlyCys: 1.974 ± 0.642
4.343GlyAsp: 4.343 ± 0.862
5.132GlyGlu: 5.132 ± 1.12
1.579GlyPhe: 1.579 ± 0.619
5.527GlyGly: 5.527 ± 1.359
1.579GlyHis: 1.579 ± 0.455
3.948GlyIle: 3.948 ± 1.98
3.158GlyLys: 3.158 ± 1.175
4.737GlyLeu: 4.737 ± 0.572
0.0GlyMet: 0.0 ± 0.0
3.948GlyAsn: 3.948 ± 0.872
4.737GlyPro: 4.737 ± 1.776
1.974GlyGln: 1.974 ± 0.774
7.501GlyArg: 7.501 ± 1.762
6.317GlySer: 6.317 ± 1.329
5.527GlyThr: 5.527 ± 0.588
4.343GlyVal: 4.343 ± 0.804
0.0GlyTrp: 0.0 ± 0.0
0.79GlyTyr: 0.79 ± 0.316
0.0GlyXaa: 0.0 ± 0.0
His
1.184HisAla: 1.184 ± 0.699
0.79HisCys: 0.79 ± 0.427
0.79HisAsp: 0.79 ± 0.499
0.79HisGlu: 0.79 ± 0.705
1.184HisPhe: 1.184 ± 0.626
1.579HisGly: 1.579 ± 0.455
0.0HisHis: 0.0 ± 0.0
1.579HisIle: 1.579 ± 0.573
0.395HisLys: 0.395 ± 0.352
1.184HisLeu: 1.184 ± 0.724
0.395HisMet: 0.395 ± 0.352
0.0HisAsn: 0.0 ± 0.0
0.395HisPro: 0.395 ± 0.348
0.79HisGln: 0.79 ± 0.427
0.395HisArg: 0.395 ± 0.275
1.579HisSer: 1.579 ± 0.496
1.579HisThr: 1.579 ± 0.619
0.0HisVal: 0.0 ± 0.0
0.79HisTrp: 0.79 ± 0.369
1.579HisTyr: 1.579 ± 0.99
0.0HisXaa: 0.0 ± 0.0
Ile
1.974IleAla: 1.974 ± 0.657
2.369IleCys: 2.369 ± 0.775
3.158IleAsp: 3.158 ± 1.171
3.948IleGlu: 3.948 ± 1.398
1.579IlePhe: 1.579 ± 0.455
4.737IleGly: 4.737 ± 2.071
0.395IleHis: 0.395 ± 0.352
1.579IleIle: 1.579 ± 0.619
1.974IleLys: 1.974 ± 0.38
3.158IleLeu: 3.158 ± 0.682
0.395IleMet: 0.395 ± 0.428
0.79IleAsn: 0.79 ± 0.336
2.764IlePro: 2.764 ± 1.452
1.184IleGln: 1.184 ± 0.547
2.764IleArg: 2.764 ± 0.955
3.158IleSer: 3.158 ± 0.991
2.369IleThr: 2.369 ± 1.08
2.764IleVal: 2.764 ± 0.841
0.0IleTrp: 0.0 ± 0.0
1.974IleTyr: 1.974 ± 0.639
0.0IleXaa: 0.0 ± 0.0
Lys
2.764LysAla: 2.764 ± 0.865
1.974LysCys: 1.974 ± 0.995
0.79LysAsp: 0.79 ± 0.336
3.553LysGlu: 3.553 ± 0.754
4.343LysPhe: 4.343 ± 1.05
1.579LysGly: 1.579 ± 0.556
0.79LysHis: 0.79 ± 0.427
0.395LysIle: 0.395 ± 0.352
2.369LysLys: 2.369 ± 1.265
3.158LysLeu: 3.158 ± 1.679
0.395LysMet: 0.395 ± 0.32
2.369LysAsn: 2.369 ± 0.597
1.974LysPro: 1.974 ± 0.764
2.369LysGln: 2.369 ± 0.598
3.553LysArg: 3.553 ± 0.468
4.737LysSer: 4.737 ± 2.339
2.369LysThr: 2.369 ± 0.724
1.974LysVal: 1.974 ± 1.099
0.0LysTrp: 0.0 ± 0.0
3.948LysTyr: 3.948 ± 2.142
0.0LysXaa: 0.0 ± 0.0
Leu
5.132LeuAla: 5.132 ± 1.764
3.553LeuCys: 3.553 ± 1.514
3.948LeuAsp: 3.948 ± 1.275
3.553LeuGlu: 3.553 ± 1.244
3.948LeuPhe: 3.948 ± 0.282
9.08LeuGly: 9.08 ± 3.293
1.184LeuHis: 1.184 ± 0.597
2.369LeuIle: 2.369 ± 0.835
5.132LeuLys: 5.132 ± 1.411
5.132LeuLeu: 5.132 ± 1.276
1.184LeuMet: 1.184 ± 0.567
1.579LeuAsn: 1.579 ± 0.395
3.948LeuPro: 3.948 ± 0.936
5.132LeuGln: 5.132 ± 1.184
4.343LeuArg: 4.343 ± 0.865
7.106LeuSer: 7.106 ± 1.411
5.527LeuThr: 5.527 ± 2.289
4.737LeuVal: 4.737 ± 0.849
1.579LeuTrp: 1.579 ± 0.395
2.764LeuTyr: 2.764 ± 0.455
0.0LeuXaa: 0.0 ± 0.0
Met
1.579MetAla: 1.579 ± 0.672
0.0MetCys: 0.0 ± 0.0
1.184MetAsp: 1.184 ± 0.842
1.184MetGlu: 1.184 ± 0.451
1.579MetPhe: 1.579 ± 0.53
1.184MetGly: 1.184 ± 0.858
0.79MetHis: 0.79 ± 0.705
1.579MetIle: 1.579 ± 0.855
0.395MetLys: 0.395 ± 0.352
1.579MetLeu: 1.579 ± 1.024
0.79MetMet: 0.79 ± 1.032
0.79MetAsn: 0.79 ± 0.336
0.0MetPro: 0.0 ± 0.0
0.79MetGln: 0.79 ± 0.427
0.0MetArg: 0.0 ± 0.0
1.184MetSer: 1.184 ± 0.626
0.0MetThr: 0.0 ± 0.0
0.79MetVal: 0.79 ± 0.705
0.0MetTrp: 0.0 ± 0.0
0.79MetTyr: 0.79 ± 0.45
0.0MetXaa: 0.0 ± 0.0
Asn
1.974AsnAla: 1.974 ± 0.558
0.79AsnCys: 0.79 ± 0.499
1.579AsnAsp: 1.579 ± 0.495
2.369AsnGlu: 2.369 ± 1.759
0.79AsnPhe: 0.79 ± 0.336
0.79AsnGly: 0.79 ± 0.336
0.0AsnHis: 0.0 ± 0.0
1.579AsnIle: 1.579 ± 0.723
1.579AsnLys: 1.579 ± 0.666
1.184AsnLeu: 1.184 ± 0.96
0.395AsnMet: 0.395 ± 0.32
1.184AsnAsn: 1.184 ± 0.699
4.737AsnPro: 4.737 ± 1.117
2.764AsnGln: 2.764 ± 0.714
3.553AsnArg: 3.553 ± 1.595
3.948AsnSer: 3.948 ± 1.244
2.369AsnThr: 2.369 ± 0.72
1.974AsnVal: 1.974 ± 0.779
0.79AsnTrp: 0.79 ± 0.336
0.395AsnTyr: 0.395 ± 0.32
0.0AsnXaa: 0.0 ± 0.0
Pro
7.896ProAla: 7.896 ± 2.094
1.579ProCys: 1.579 ± 0.655
3.948ProAsp: 3.948 ± 1.658
4.737ProGlu: 4.737 ± 1.406
1.184ProPhe: 1.184 ± 0.733
5.922ProGly: 5.922 ± 2.918
0.0ProHis: 0.0 ± 0.0
3.948ProIle: 3.948 ± 0.567
2.764ProLys: 2.764 ± 0.848
7.106ProLeu: 7.106 ± 1.741
0.395ProMet: 0.395 ± 0.32
1.579ProAsn: 1.579 ± 0.934
10.659ProPro: 10.659 ± 2.65
2.369ProGln: 2.369 ± 1.209
3.158ProArg: 3.158 ± 1.102
4.737ProSer: 4.737 ± 1.003
4.737ProThr: 4.737 ± 1.443
3.948ProVal: 3.948 ± 1.65
1.184ProTrp: 1.184 ± 0.553
2.369ProTyr: 2.369 ± 0.953
0.0ProXaa: 0.0 ± 0.0
Gln
3.948GlnAla: 3.948 ± 1.154
0.79GlnCys: 0.79 ± 0.427
1.184GlnAsp: 1.184 ± 0.699
1.579GlnGlu: 1.579 ± 0.855
0.79GlnPhe: 0.79 ± 0.705
2.764GlnGly: 2.764 ± 0.829
1.184GlnHis: 1.184 ± 0.36
1.184GlnIle: 1.184 ± 0.48
1.184GlnLys: 1.184 ± 0.733
4.343GlnLeu: 4.343 ± 0.974
1.974GlnMet: 1.974 ± 0.561
1.184GlnAsn: 1.184 ± 0.316
1.579GlnPro: 1.579 ± 0.455
7.106GlnGln: 7.106 ± 2.847
4.343GlnArg: 4.343 ± 1.271
3.553GlnSer: 3.553 ± 1.364
3.553GlnThr: 3.553 ± 0.702
2.369GlnVal: 2.369 ± 0.712
1.184GlnTrp: 1.184 ± 0.627
0.79GlnTyr: 0.79 ± 0.45
0.0GlnXaa: 0.0 ± 0.0
Arg
2.764ArgAla: 2.764 ± 0.782
2.764ArgCys: 2.764 ± 1.072
3.948ArgAsp: 3.948 ± 1.963
3.948ArgGlu: 3.948 ± 0.946
2.764ArgPhe: 2.764 ± 0.913
3.158ArgGly: 3.158 ± 1.224
1.579ArgHis: 1.579 ± 0.739
3.158ArgIle: 3.158 ± 0.964
5.527ArgLys: 5.527 ± 1.001
3.948ArgLeu: 3.948 ± 1.112
0.79ArgMet: 0.79 ± 0.427
1.184ArgAsn: 1.184 ± 0.627
5.922ArgPro: 5.922 ± 1.701
3.553ArgGln: 3.553 ± 1.375
9.08ArgArg: 9.08 ± 2.61
4.343ArgSer: 4.343 ± 1.491
5.132ArgThr: 5.132 ± 0.823
4.737ArgVal: 4.737 ± 0.617
1.579ArgTrp: 1.579 ± 0.855
3.158ArgTyr: 3.158 ± 1.317
0.0ArgXaa: 0.0 ± 0.0
Ser
2.369SerAla: 2.369 ± 1.087
1.974SerCys: 1.974 ± 1.148
7.106SerAsp: 7.106 ± 1.3
3.948SerGlu: 3.948 ± 0.925
3.948SerPhe: 3.948 ± 1.698
7.106SerGly: 7.106 ± 0.97
1.579SerHis: 1.579 ± 0.727
4.343SerIle: 4.343 ± 1.18
0.79SerLys: 0.79 ± 0.369
7.501SerLeu: 7.501 ± 1.061
1.184SerMet: 1.184 ± 1.057
4.343SerAsn: 4.343 ± 2.391
4.737SerPro: 4.737 ± 2.78
3.948SerGln: 3.948 ± 1.211
7.501SerArg: 7.501 ± 2.655
7.106SerSer: 7.106 ± 2.178
7.896SerThr: 7.896 ± 1.553
3.948SerVal: 3.948 ± 1.045
0.79SerTrp: 0.79 ± 0.427
0.79SerTyr: 0.79 ± 0.705
0.0SerXaa: 0.0 ± 0.0
Thr
1.579ThrAla: 1.579 ± 1.068
0.79ThrCys: 0.79 ± 0.363
4.343ThrAsp: 4.343 ± 1.547
2.764ThrGlu: 2.764 ± 0.882
2.764ThrPhe: 2.764 ± 0.804
8.291ThrGly: 8.291 ± 2.185
1.579ThrHis: 1.579 ± 0.573
3.158ThrIle: 3.158 ± 2.424
3.158ThrLys: 3.158 ± 0.99
3.948ThrLeu: 3.948 ± 1.167
1.579ThrMet: 1.579 ± 1.41
2.369ThrAsn: 2.369 ± 0.746
6.317ThrPro: 6.317 ± 1.47
1.974ThrGln: 1.974 ± 0.387
7.896ThrArg: 7.896 ± 1.728
5.922ThrSer: 5.922 ± 1.309
3.553ThrThr: 3.553 ± 1.839
7.501ThrVal: 7.501 ± 1.62
0.395ThrTrp: 0.395 ± 0.275
0.79ThrTyr: 0.79 ± 0.656
0.0ThrXaa: 0.0 ± 0.0
Val
3.553ValAla: 3.553 ± 0.902
0.79ValCys: 0.79 ± 0.705
4.737ValAsp: 4.737 ± 0.901
1.974ValGlu: 1.974 ± 0.855
2.764ValPhe: 2.764 ± 0.941
2.764ValGly: 2.764 ± 0.402
1.184ValHis: 1.184 ± 0.618
2.369ValIle: 2.369 ± 1.717
2.369ValLys: 2.369 ± 0.631
5.527ValLeu: 5.527 ± 1.387
0.79ValMet: 0.79 ± 0.336
1.184ValAsn: 1.184 ± 0.699
5.132ValPro: 5.132 ± 0.79
3.158ValGln: 3.158 ± 0.694
3.158ValArg: 3.158 ± 1.033
6.317ValSer: 6.317 ± 0.971
5.922ValThr: 5.922 ± 1.249
4.737ValVal: 4.737 ± 0.813
1.184ValTrp: 1.184 ± 0.567
0.395ValTyr: 0.395 ± 0.459
0.0ValXaa: 0.0 ± 0.0
Trp
1.184TrpAla: 1.184 ± 0.36
0.0TrpCys: 0.0 ± 0.0
1.184TrpAsp: 1.184 ± 0.627
0.79TrpGlu: 0.79 ± 0.369
0.395TrpPhe: 0.395 ± 0.352
0.79TrpGly: 0.79 ± 0.369
0.0TrpHis: 0.0 ± 0.0
0.79TrpIle: 0.79 ± 0.705
1.184TrpLys: 1.184 ± 0.732
1.974TrpLeu: 1.974 ± 0.639
0.0TrpMet: 0.0 ± 0.0
0.79TrpAsn: 0.79 ± 0.64
0.395TrpPro: 0.395 ± 0.352
0.395TrpGln: 0.395 ± 0.352
0.395TrpArg: 0.395 ± 0.275
0.395TrpSer: 0.395 ± 0.275
0.395TrpThr: 0.395 ± 0.275
1.184TrpVal: 1.184 ± 0.451
0.0TrpTrp: 0.0 ± 0.0
1.184TrpTyr: 1.184 ± 0.826
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.369TyrAla: 2.369 ± 0.72
0.0TyrCys: 0.0 ± 0.0
1.579TyrAsp: 1.579 ± 0.53
2.369TyrGlu: 2.369 ± 0.51
1.184TyrPhe: 1.184 ± 0.96
1.974TyrGly: 1.974 ± 0.991
1.579TyrHis: 1.579 ± 0.592
0.395TyrIle: 0.395 ± 0.275
1.579TyrLys: 1.579 ± 0.723
2.369TyrLeu: 2.369 ± 0.933
0.79TyrMet: 0.79 ± 0.705
0.79TyrAsn: 0.79 ± 0.64
1.579TyrPro: 1.579 ± 0.619
1.184TyrGln: 1.184 ± 0.61
3.553TyrArg: 3.553 ± 0.908
1.184TyrSer: 1.184 ± 0.605
0.79TyrThr: 0.79 ± 0.45
1.974TyrVal: 1.974 ± 0.471
1.184TyrTrp: 1.184 ± 0.36
1.974TyrTyr: 1.974 ± 0.722
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 6 proteins (2534 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski