Amino acid dipepetide frequency for Rice yellow stunt virus (RYSV) (Rice transitory yellowing virus)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
3.635AlaAla: 3.635 ± 0.665
1.212AlaCys: 1.212 ± 0.241
3.877AlaAsp: 3.877 ± 0.842
4.119AlaGlu: 4.119 ± 0.846
0.727AlaPhe: 0.727 ± 0.302
3.635AlaGly: 3.635 ± 1.192
0.969AlaHis: 0.969 ± 0.543
3.877AlaIle: 3.877 ± 0.701
3.15AlaLys: 3.15 ± 2.11
4.362AlaLeu: 4.362 ± 0.664
0.969AlaMet: 0.969 ± 0.779
2.181AlaAsn: 2.181 ± 1.408
1.696AlaPro: 1.696 ± 0.802
1.938AlaGln: 1.938 ± 1.233
3.15AlaArg: 3.15 ± 1.136
3.15AlaSer: 3.15 ± 1.582
4.119AlaThr: 4.119 ± 0.858
2.423AlaVal: 2.423 ± 0.377
1.454AlaTrp: 1.454 ± 0.553
2.181AlaTyr: 2.181 ± 0.703
0.0AlaXaa: 0.0 ± 0.0
Cys
0.727CysAla: 0.727 ± 0.381
0.485CysCys: 0.485 ± 0.271
1.938CysAsp: 1.938 ± 0.704
0.727CysGlu: 0.727 ± 0.336
0.242CysPhe: 0.242 ± 0.147
0.0CysGly: 0.0 ± 0.0
0.242CysHis: 0.242 ± 0.147
1.696CysIle: 1.696 ± 0.305
1.938CysLys: 1.938 ± 0.821
0.485CysLeu: 0.485 ± 0.293
0.485CysMet: 0.485 ± 0.213
1.212CysAsn: 1.212 ± 0.333
0.485CysPro: 0.485 ± 0.458
0.969CysGln: 0.969 ± 0.211
0.485CysArg: 0.485 ± 0.271
2.423CysSer: 2.423 ± 0.979
1.212CysThr: 1.212 ± 0.444
0.727CysVal: 0.727 ± 0.449
0.242CysTrp: 0.242 ± 0.147
1.454CysTyr: 1.454 ± 0.627
0.0CysXaa: 0.0 ± 0.0
Asp
3.392AspAla: 3.392 ± 1.264
0.969AspCys: 0.969 ± 0.363
3.15AspAsp: 3.15 ± 0.837
5.331AspGlu: 5.331 ± 1.437
1.696AspPhe: 1.696 ± 0.723
3.635AspGly: 3.635 ± 1.454
1.212AspHis: 1.212 ± 0.582
5.331AspIle: 5.331 ± 1.585
3.635AspLys: 3.635 ± 0.866
5.573AspLeu: 5.573 ± 0.69
1.696AspMet: 1.696 ± 0.587
2.908AspAsn: 2.908 ± 0.585
3.392AspPro: 3.392 ± 1.112
0.969AspGln: 0.969 ± 0.79
2.181AspArg: 2.181 ± 0.521
2.423AspSer: 2.423 ± 0.539
4.362AspThr: 4.362 ± 0.909
4.846AspVal: 4.846 ± 1.204
0.969AspTrp: 0.969 ± 0.587
3.392AspTyr: 3.392 ± 0.855
0.0AspXaa: 0.0 ± 0.0
Glu
4.604GluAla: 4.604 ± 1.113
1.696GluCys: 1.696 ± 0.432
4.362GluAsp: 4.362 ± 1.275
3.15GluGlu: 3.15 ± 0.833
0.969GluPhe: 0.969 ± 0.46
5.331GluGly: 5.331 ± 0.809
1.696GluHis: 1.696 ± 0.503
2.665GluIle: 2.665 ± 0.736
3.635GluLys: 3.635 ± 0.896
6.058GluLeu: 6.058 ± 1.055
0.969GluMet: 0.969 ± 0.483
1.696GluAsn: 1.696 ± 0.337
1.696GluPro: 1.696 ± 0.454
1.454GluGln: 1.454 ± 0.349
2.908GluArg: 2.908 ± 0.668
4.604GluSer: 4.604 ± 0.787
4.604GluThr: 4.604 ± 1.342
3.15GluVal: 3.15 ± 1.113
0.485GluTrp: 0.485 ± 0.262
0.485GluTyr: 0.485 ± 0.302
0.0GluXaa: 0.0 ± 0.0
Phe
1.212PheAla: 1.212 ± 0.468
0.242PheCys: 0.242 ± 0.314
1.938PheAsp: 1.938 ± 0.431
1.454PheGlu: 1.454 ± 0.458
1.454PhePhe: 1.454 ± 0.638
1.212PheGly: 1.212 ± 0.442
1.454PheHis: 1.454 ± 0.672
1.938PheIle: 1.938 ± 0.687
2.181PheLys: 2.181 ± 0.54
2.423PheLeu: 2.423 ± 0.826
0.969PheMet: 0.969 ± 0.421
0.969PheAsn: 0.969 ± 0.363
1.696PhePro: 1.696 ± 0.765
0.727PheGln: 0.727 ± 0.569
1.212PheArg: 1.212 ± 0.49
4.119PheSer: 4.119 ± 0.857
1.212PheThr: 1.212 ± 0.451
1.696PheVal: 1.696 ± 0.364
0.969PheTrp: 0.969 ± 0.425
0.727PheTyr: 0.727 ± 0.44
0.0PheXaa: 0.0 ± 0.0
Gly
4.846GlyAla: 4.846 ± 1.045
0.485GlyCys: 0.485 ± 0.213
2.908GlyAsp: 2.908 ± 1.079
2.665GlyGlu: 2.665 ± 0.515
1.938GlyPhe: 1.938 ± 0.41
6.058GlyGly: 6.058 ± 1.647
1.454GlyHis: 1.454 ± 0.49
2.908GlyIle: 2.908 ± 0.86
4.119GlyLys: 4.119 ± 0.9
5.088GlyLeu: 5.088 ± 1.155
1.212GlyMet: 1.212 ± 0.763
3.877GlyAsn: 3.877 ± 0.766
2.665GlyPro: 2.665 ± 0.656
1.696GlyGln: 1.696 ± 0.857
2.423GlyArg: 2.423 ± 0.926
4.362GlySer: 4.362 ± 0.894
2.908GlyThr: 2.908 ± 0.785
5.331GlyVal: 5.331 ± 0.934
1.696GlyTrp: 1.696 ± 0.464
1.696GlyTyr: 1.696 ± 0.656
0.0GlyXaa: 0.0 ± 0.0
His
0.727HisAla: 0.727 ± 0.322
0.0HisCys: 0.0 ± 0.0
1.454HisAsp: 1.454 ± 0.906
0.727HisGlu: 0.727 ± 0.313
1.212HisPhe: 1.212 ± 0.366
1.212HisGly: 1.212 ± 0.49
1.212HisHis: 1.212 ± 0.551
2.665HisIle: 2.665 ± 1.113
1.212HisLys: 1.212 ± 0.366
3.635HisLeu: 3.635 ± 0.606
0.727HisMet: 0.727 ± 0.302
0.485HisAsn: 0.485 ± 0.271
1.696HisPro: 1.696 ± 0.539
1.696HisGln: 1.696 ± 0.597
0.969HisArg: 0.969 ± 0.543
0.727HisSer: 0.727 ± 0.569
1.454HisThr: 1.454 ± 0.462
0.969HisVal: 0.969 ± 0.425
0.242HisTrp: 0.242 ± 0.314
0.969HisTyr: 0.969 ± 0.606
0.0HisXaa: 0.0 ± 0.0
Ile
3.635IleAla: 3.635 ± 1.507
2.181IleCys: 2.181 ± 0.553
4.846IleAsp: 4.846 ± 0.972
2.423IleGlu: 2.423 ± 0.773
0.969IlePhe: 0.969 ± 0.587
3.635IleGly: 3.635 ± 1.355
1.454IleHis: 1.454 ± 0.675
7.269IleIle: 7.269 ± 1.276
4.604IleLys: 4.604 ± 1.114
6.785IleLeu: 6.785 ± 1.72
2.665IleMet: 2.665 ± 0.743
3.635IleAsn: 3.635 ± 0.976
3.15IlePro: 3.15 ± 0.925
2.423IleGln: 2.423 ± 0.339
3.392IleArg: 3.392 ± 0.917
7.027IleSer: 7.027 ± 1.186
6.542IleThr: 6.542 ± 1.339
3.877IleVal: 3.877 ± 0.795
1.212IleTrp: 1.212 ± 0.374
2.908IleTyr: 2.908 ± 0.948
0.0IleXaa: 0.0 ± 0.0
Lys
2.665LysAla: 2.665 ± 0.874
0.242LysCys: 0.242 ± 0.147
2.665LysAsp: 2.665 ± 0.811
5.331LysGlu: 5.331 ± 0.834
1.938LysPhe: 1.938 ± 0.413
3.635LysGly: 3.635 ± 0.682
1.454LysHis: 1.454 ± 0.522
5.331LysIle: 5.331 ± 0.863
2.665LysLys: 2.665 ± 0.639
6.3LysLeu: 6.3 ± 1.279
2.665LysMet: 2.665 ± 0.584
2.181LysAsn: 2.181 ± 0.798
2.423LysPro: 2.423 ± 0.383
1.938LysGln: 1.938 ± 0.71
4.604LysArg: 4.604 ± 1.193
4.604LysSer: 4.604 ± 0.624
4.604LysThr: 4.604 ± 0.918
3.635LysVal: 3.635 ± 1.17
0.485LysTrp: 0.485 ± 0.213
2.665LysTyr: 2.665 ± 1.141
0.0LysXaa: 0.0 ± 0.0
Leu
4.846LeuAla: 4.846 ± 1.793
1.454LeuCys: 1.454 ± 0.616
4.119LeuAsp: 4.119 ± 1.25
4.846LeuGlu: 4.846 ± 0.807
3.392LeuPhe: 3.392 ± 0.633
6.3LeuGly: 6.3 ± 1.339
2.181LeuHis: 2.181 ± 1.07
7.027LeuIle: 7.027 ± 1.908
7.269LeuLys: 7.269 ± 1.519
6.3LeuLeu: 6.3 ± 1.163
1.938LeuMet: 1.938 ± 0.623
4.362LeuAsn: 4.362 ± 0.463
3.877LeuPro: 3.877 ± 1.051
2.423LeuGln: 2.423 ± 0.645
4.119LeuArg: 4.119 ± 0.765
8.238LeuSer: 8.238 ± 0.631
6.542LeuThr: 6.542 ± 1.1
5.573LeuVal: 5.573 ± 0.837
0.485LeuTrp: 0.485 ± 0.213
4.362LeuTyr: 4.362 ± 0.687
0.0LeuXaa: 0.0 ± 0.0
Met
2.181MetAla: 2.181 ± 0.401
0.485MetCys: 0.485 ± 0.213
1.938MetAsp: 1.938 ± 0.664
1.696MetGlu: 1.696 ± 0.992
0.727MetPhe: 0.727 ± 0.302
2.181MetGly: 2.181 ± 0.54
0.485MetHis: 0.485 ± 0.302
0.727MetIle: 0.727 ± 0.259
1.696MetLys: 1.696 ± 0.546
2.181MetLeu: 2.181 ± 0.644
0.969MetMet: 0.969 ± 0.368
0.727MetAsn: 0.727 ± 0.259
0.969MetPro: 0.969 ± 0.391
0.485MetGln: 0.485 ± 0.293
2.908MetArg: 2.908 ± 0.606
2.423MetSer: 2.423 ± 0.473
2.665MetThr: 2.665 ± 0.402
1.938MetVal: 1.938 ± 0.602
0.0MetTrp: 0.0 ± 0.0
2.665MetTyr: 2.665 ± 0.568
0.0MetXaa: 0.0 ± 0.0
Asn
2.423AsnAla: 2.423 ± 0.879
1.212AsnCys: 1.212 ± 0.442
2.665AsnAsp: 2.665 ± 0.776
1.454AsnGlu: 1.454 ± 0.325
0.727AsnPhe: 0.727 ± 0.336
1.212AsnGly: 1.212 ± 0.801
1.454AsnHis: 1.454 ± 0.448
4.604AsnIle: 4.604 ± 1.105
3.392AsnLys: 3.392 ± 1.001
4.604AsnLeu: 4.604 ± 0.934
1.212AsnMet: 1.212 ± 0.582
4.362AsnAsn: 4.362 ± 1.016
2.181AsnPro: 2.181 ± 0.767
1.938AsnGln: 1.938 ± 0.719
1.454AsnArg: 1.454 ± 0.787
3.15AsnSer: 3.15 ± 1.235
3.877AsnThr: 3.877 ± 0.991
3.877AsnVal: 3.877 ± 1.174
1.212AsnTrp: 1.212 ± 0.315
2.181AsnTyr: 2.181 ± 0.691
0.0AsnXaa: 0.0 ± 0.0
Pro
1.212ProAla: 1.212 ± 0.677
0.485ProCys: 0.485 ± 0.213
2.908ProAsp: 2.908 ± 0.887
3.635ProGlu: 3.635 ± 0.608
1.938ProPhe: 1.938 ± 0.392
1.454ProGly: 1.454 ± 0.634
0.727ProHis: 0.727 ± 0.302
3.15ProIle: 3.15 ± 1.567
1.696ProLys: 1.696 ± 0.549
4.846ProLeu: 4.846 ± 0.677
1.454ProMet: 1.454 ± 0.673
2.423ProAsn: 2.423 ± 1.308
2.908ProPro: 2.908 ± 1.113
2.181ProGln: 2.181 ± 1.053
2.908ProArg: 2.908 ± 0.655
2.665ProSer: 2.665 ± 1.041
2.423ProThr: 2.423 ± 0.552
2.665ProVal: 2.665 ± 0.528
0.485ProTrp: 0.485 ± 0.37
1.696ProTyr: 1.696 ± 0.3
0.0ProXaa: 0.0 ± 0.0
Gln
0.969GlnAla: 0.969 ± 0.62
0.727GlnCys: 0.727 ± 0.302
3.15GlnAsp: 3.15 ± 0.458
2.908GlnGlu: 2.908 ± 1.122
0.969GlnPhe: 0.969 ± 0.34
1.938GlnGly: 1.938 ± 0.714
1.212GlnHis: 1.212 ± 0.468
3.15GlnIle: 3.15 ± 0.402
1.696GlnLys: 1.696 ± 0.521
1.696GlnLeu: 1.696 ± 0.57
1.696GlnMet: 1.696 ± 0.607
1.454GlnAsn: 1.454 ± 0.54
0.969GlnPro: 0.969 ± 0.738
2.665GlnGln: 2.665 ± 1.055
1.212GlnArg: 1.212 ± 0.25
1.938GlnSer: 1.938 ± 0.68
1.938GlnThr: 1.938 ± 0.86
1.696GlnVal: 1.696 ± 0.599
0.0GlnTrp: 0.0 ± 0.0
1.212GlnTyr: 1.212 ± 1.152
0.0GlnXaa: 0.0 ± 0.0
Arg
2.665ArgAla: 2.665 ± 0.658
0.485ArgCys: 0.485 ± 0.293
2.908ArgAsp: 2.908 ± 0.689
2.665ArgGlu: 2.665 ± 0.555
1.212ArgPhe: 1.212 ± 0.555
2.908ArgGly: 2.908 ± 0.873
0.727ArgHis: 0.727 ± 0.302
4.604ArgIle: 4.604 ± 1.096
2.181ArgLys: 2.181 ± 0.583
4.362ArgLeu: 4.362 ± 0.684
2.423ArgMet: 2.423 ± 0.726
0.727ArgAsn: 0.727 ± 0.569
1.696ArgPro: 1.696 ± 0.412
1.696ArgGln: 1.696 ± 0.719
2.423ArgArg: 2.423 ± 0.782
6.3ArgSer: 6.3 ± 1.266
2.181ArgThr: 2.181 ± 0.847
4.604ArgVal: 4.604 ± 0.29
0.242ArgTrp: 0.242 ± 0.258
1.696ArgTyr: 1.696 ± 0.593
0.0ArgXaa: 0.0 ± 0.0
Ser
3.635SerAla: 3.635 ± 1.289
0.727SerCys: 0.727 ± 0.773
4.119SerAsp: 4.119 ± 0.789
4.362SerGlu: 4.362 ± 0.424
2.665SerPhe: 2.665 ± 0.788
5.815SerGly: 5.815 ± 1.576
1.938SerHis: 1.938 ± 0.547
4.362SerIle: 4.362 ± 0.519
6.058SerLys: 6.058 ± 1.23
6.058SerLeu: 6.058 ± 1.125
1.696SerMet: 1.696 ± 0.793
5.088SerAsn: 5.088 ± 1.367
3.635SerPro: 3.635 ± 0.652
2.181SerGln: 2.181 ± 0.617
5.088SerArg: 5.088 ± 1.085
7.027SerSer: 7.027 ± 1.894
7.027SerThr: 7.027 ± 0.876
2.908SerVal: 2.908 ± 0.934
2.181SerTrp: 2.181 ± 0.513
4.119SerTyr: 4.119 ± 1.665
0.0SerXaa: 0.0 ± 0.0
Thr
2.665ThrAla: 2.665 ± 0.798
1.212ThrCys: 1.212 ± 0.25
5.088ThrAsp: 5.088 ± 1.556
4.119ThrGlu: 4.119 ± 0.842
2.181ThrPhe: 2.181 ± 0.26
3.877ThrGly: 3.877 ± 0.943
1.938ThrHis: 1.938 ± 0.66
3.15ThrIle: 3.15 ± 0.809
2.423ThrLys: 2.423 ± 0.949
6.542ThrLeu: 6.542 ± 1.409
3.392ThrMet: 3.392 ± 1.035
3.392ThrAsn: 3.392 ± 1.033
4.846ThrPro: 4.846 ± 0.361
3.15ThrGln: 3.15 ± 1.166
1.938ThrArg: 1.938 ± 0.341
5.815ThrSer: 5.815 ± 1.173
4.362ThrThr: 4.362 ± 1.237
5.573ThrVal: 5.573 ± 0.882
1.454ThrTrp: 1.454 ± 0.814
2.665ThrTyr: 2.665 ± 0.399
0.0ThrXaa: 0.0 ± 0.0
Val
4.119ValAla: 4.119 ± 1.129
1.696ValCys: 1.696 ± 0.614
3.877ValAsp: 3.877 ± 0.388
2.908ValGlu: 2.908 ± 0.472
2.181ValPhe: 2.181 ± 0.344
3.635ValGly: 3.635 ± 0.848
1.212ValHis: 1.212 ± 0.518
5.088ValIle: 5.088 ± 0.941
4.604ValLys: 4.604 ± 1.211
5.573ValLeu: 5.573 ± 1.139
2.423ValMet: 2.423 ± 0.624
3.877ValAsn: 3.877 ± 0.887
1.696ValPro: 1.696 ± 0.36
1.454ValGln: 1.454 ± 0.713
2.665ValArg: 2.665 ± 0.889
3.635ValSer: 3.635 ± 1.07
4.846ValThr: 4.846 ± 0.889
2.665ValVal: 2.665 ± 0.786
0.485ValTrp: 0.485 ± 0.515
1.454ValTyr: 1.454 ± 0.448
0.0ValXaa: 0.0 ± 0.0
Trp
1.212TrpAla: 1.212 ± 0.468
0.242TrpCys: 0.242 ± 0.147
0.727TrpAsp: 0.727 ± 0.316
0.485TrpGlu: 0.485 ± 0.295
0.485TrpPhe: 0.485 ± 0.213
0.969TrpGly: 0.969 ± 0.587
0.0TrpHis: 0.0 ± 0.0
1.454TrpIle: 1.454 ± 0.49
1.454TrpLys: 1.454 ± 0.441
1.938TrpLeu: 1.938 ± 0.458
0.0TrpMet: 0.0 ± 0.0
0.727TrpAsn: 0.727 ± 0.302
0.485TrpPro: 0.485 ± 0.262
0.242TrpGln: 0.242 ± 0.147
1.212TrpArg: 1.212 ± 0.678
0.727TrpSer: 0.727 ± 0.336
0.969TrpThr: 0.969 ± 0.39
0.727TrpVal: 0.727 ± 0.449
0.485TrpTrp: 0.485 ± 0.213
0.969TrpTyr: 0.969 ± 0.496
0.0TrpXaa: 0.0 ± 0.0
Tyr
1.454TyrAla: 1.454 ± 0.633
1.938TyrCys: 1.938 ± 0.992
2.423TyrAsp: 2.423 ± 0.858
1.454TyrGlu: 1.454 ± 0.66
2.181TyrPhe: 2.181 ± 0.777
1.696TyrGly: 1.696 ± 0.601
0.969TyrHis: 0.969 ± 0.604
3.635TyrIle: 3.635 ± 0.576
2.423TyrLys: 2.423 ± 0.604
4.846TyrLeu: 4.846 ± 0.792
0.242TyrMet: 0.242 ± 0.336
2.908TyrAsn: 2.908 ± 0.695
1.696TyrPro: 1.696 ± 1.081
0.969TyrGln: 0.969 ± 0.587
1.454TyrArg: 1.454 ± 0.289
5.331TyrSer: 5.331 ± 0.787
1.938TyrThr: 1.938 ± 0.811
1.212TyrVal: 1.212 ± 0.357
0.727TyrTrp: 0.727 ± 0.302
0.727TyrTyr: 0.727 ± 0.449
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (4128 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski