Amino acid dipepetide frequency for Barley yellow striate mosaic cytorhabdovirus

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
4.055AlaAla: 4.055 ± 1.928
0.507AlaCys: 0.507 ± 0.361
4.815AlaAsp: 4.815 ± 1.784
3.041AlaGlu: 3.041 ± 1.18
2.027AlaPhe: 2.027 ± 0.706
2.534AlaGly: 2.534 ± 0.935
0.76AlaHis: 0.76 ± 0.336
3.041AlaIle: 3.041 ± 1.186
4.562AlaLys: 4.562 ± 0.775
4.055AlaLeu: 4.055 ± 0.927
0.507AlaMet: 0.507 ± 0.309
2.534AlaAsn: 2.534 ± 1.183
1.774AlaPro: 1.774 ± 0.658
1.014AlaGln: 1.014 ± 0.403
3.294AlaArg: 3.294 ± 0.706
5.322AlaSer: 5.322 ± 0.842
3.548AlaThr: 3.548 ± 1.236
2.534AlaVal: 2.534 ± 1.058
0.507AlaTrp: 0.507 ± 0.433
2.281AlaTyr: 2.281 ± 0.798
0.0AlaXaa: 0.0 ± 0.0
Cys
0.253CysAla: 0.253 ± 0.148
0.507CysCys: 0.507 ± 0.667
1.521CysAsp: 1.521 ± 0.874
0.0CysGlu: 0.0 ± 0.0
0.253CysPhe: 0.253 ± 0.405
0.507CysGly: 0.507 ± 0.309
0.507CysHis: 0.507 ± 0.477
1.774CysIle: 1.774 ± 0.413
1.267CysLys: 1.267 ± 0.605
1.267CysLeu: 1.267 ± 0.759
0.253CysMet: 0.253 ± 0.148
1.014CysAsn: 1.014 ± 1.028
2.534CysPro: 2.534 ± 0.738
0.507CysGln: 0.507 ± 0.297
0.76CysArg: 0.76 ± 0.856
1.014CysSer: 1.014 ± 0.722
0.507CysThr: 0.507 ± 0.737
0.76CysVal: 0.76 ± 0.445
0.253CysTrp: 0.253 ± 0.148
0.507CysTyr: 0.507 ± 0.309
0.0CysXaa: 0.0 ± 0.0
Asp
2.281AspAla: 2.281 ± 0.694
1.774AspCys: 1.774 ± 0.769
2.534AspAsp: 2.534 ± 1.321
3.041AspGlu: 3.041 ± 1.225
1.774AspPhe: 1.774 ± 0.614
1.774AspGly: 1.774 ± 0.717
1.014AspHis: 1.014 ± 0.402
4.562AspIle: 4.562 ± 0.697
4.055AspLys: 4.055 ± 0.723
5.322AspLeu: 5.322 ± 1.544
1.774AspMet: 1.774 ± 0.394
4.308AspAsn: 4.308 ± 1.392
4.055AspPro: 4.055 ± 1.11
2.027AspGln: 2.027 ± 0.59
3.801AspArg: 3.801 ± 1.483
3.801AspSer: 3.801 ± 0.912
2.027AspThr: 2.027 ± 0.944
4.562AspVal: 4.562 ± 1.484
0.76AspTrp: 0.76 ± 0.314
1.267AspTyr: 1.267 ± 0.683
0.0AspXaa: 0.0 ± 0.0
Glu
2.281GluAla: 2.281 ± 0.745
0.507GluCys: 0.507 ± 0.309
4.308GluAsp: 4.308 ± 1.363
4.562GluGlu: 4.562 ± 2.253
2.281GluPhe: 2.281 ± 0.816
3.294GluGly: 3.294 ± 1.068
1.774GluHis: 1.774 ± 0.662
5.829GluIle: 5.829 ± 1.545
5.829GluLys: 5.829 ± 0.862
6.082GluLeu: 6.082 ± 1.812
2.788GluMet: 2.788 ± 0.887
3.294GluAsn: 3.294 ± 0.575
1.521GluPro: 1.521 ± 0.619
1.774GluGln: 1.774 ± 0.57
3.294GluArg: 3.294 ± 1.252
3.801GluSer: 3.801 ± 0.71
3.801GluThr: 3.801 ± 1.471
5.068GluVal: 5.068 ± 1.044
0.507GluTrp: 0.507 ± 0.589
1.521GluTyr: 1.521 ± 0.713
0.0GluXaa: 0.0 ± 0.0
Phe
1.774PheAla: 1.774 ± 1.332
0.76PheCys: 0.76 ± 0.445
2.027PheAsp: 2.027 ± 0.524
1.774PheGlu: 1.774 ± 0.436
0.76PhePhe: 0.76 ± 0.363
2.281PheGly: 2.281 ± 0.55
0.76PheHis: 0.76 ± 0.314
3.041PheIle: 3.041 ± 1.259
3.041PheLys: 3.041 ± 0.718
5.575PheLeu: 5.575 ± 1.397
0.253PheMet: 0.253 ± 0.148
0.507PheAsn: 0.507 ± 0.309
1.267PhePro: 1.267 ± 0.561
2.027PheGln: 2.027 ± 0.783
2.281PheArg: 2.281 ± 0.793
5.829PheSer: 5.829 ± 1.128
2.027PheThr: 2.027 ± 0.764
0.76PheVal: 0.76 ± 0.466
0.507PheTrp: 0.507 ± 0.507
1.267PheTyr: 1.267 ± 0.48
0.0PheXaa: 0.0 ± 0.0
Gly
2.281GlyAla: 2.281 ± 0.8
0.76GlyCys: 0.76 ± 1.105
2.534GlyAsp: 2.534 ± 0.783
6.082GlyGlu: 6.082 ± 1.374
1.267GlyPhe: 1.267 ± 0.44
2.788GlyGly: 2.788 ± 0.954
1.267GlyHis: 1.267 ± 0.499
7.603GlyIle: 7.603 ± 1.143
3.294GlyLys: 3.294 ± 1.081
6.082GlyLeu: 6.082 ± 0.971
3.041GlyMet: 3.041 ± 1.579
1.774GlyAsn: 1.774 ± 0.73
0.507GlyPro: 0.507 ± 0.333
1.014GlyGln: 1.014 ± 0.339
2.281GlyArg: 2.281 ± 0.884
3.548GlySer: 3.548 ± 1.293
1.774GlyThr: 1.774 ± 0.576
4.308GlyVal: 4.308 ± 0.582
1.521GlyTrp: 1.521 ± 0.656
2.281GlyTyr: 2.281 ± 0.687
0.0GlyXaa: 0.0 ± 0.0
His
1.267HisAla: 1.267 ± 0.456
0.507HisCys: 0.507 ± 0.309
1.267HisAsp: 1.267 ± 0.727
1.774HisGlu: 1.774 ± 0.712
0.76HisPhe: 0.76 ± 0.356
0.76HisGly: 0.76 ± 0.386
0.507HisHis: 0.507 ± 0.337
2.788HisIle: 2.788 ± 1.382
1.267HisLys: 1.267 ± 0.592
2.281HisLeu: 2.281 ± 0.849
0.0HisMet: 0.0 ± 0.0
1.267HisAsn: 1.267 ± 0.613
1.521HisPro: 1.521 ± 0.611
0.0HisGln: 0.0 ± 0.0
0.76HisArg: 0.76 ± 0.359
1.267HisSer: 1.267 ± 0.582
0.76HisThr: 0.76 ± 0.7
3.294HisVal: 3.294 ± 1.028
0.253HisTrp: 0.253 ± 0.148
0.507HisTyr: 0.507 ± 0.309
0.0HisXaa: 0.0 ± 0.0
Ile
6.336IleAla: 6.336 ± 0.965
0.253IleCys: 0.253 ± 0.281
3.294IleAsp: 3.294 ± 1.101
3.041IleGlu: 3.041 ± 0.943
4.562IlePhe: 4.562 ± 0.686
4.055IleGly: 4.055 ± 1.226
1.774IleHis: 1.774 ± 0.787
3.801IleIle: 3.801 ± 1.277
4.055IleLys: 4.055 ± 0.997
5.829IleLeu: 5.829 ± 1.793
2.027IleMet: 2.027 ± 1.012
3.294IleAsn: 3.294 ± 0.698
3.294IlePro: 3.294 ± 1.236
2.281IleGln: 2.281 ± 0.676
4.308IleArg: 4.308 ± 0.359
7.856IleSer: 7.856 ± 2.38
4.562IleThr: 4.562 ± 1.41
3.801IleVal: 3.801 ± 1.334
0.76IleTrp: 0.76 ± 0.314
2.281IleTyr: 2.281 ± 0.606
0.0IleXaa: 0.0 ± 0.0
Lys
1.521LysAla: 1.521 ± 0.606
1.521LysCys: 1.521 ± 0.614
4.308LysAsp: 4.308 ± 1.363
6.082LysGlu: 6.082 ± 0.514
2.788LysPhe: 2.788 ± 0.455
6.082LysGly: 6.082 ± 1.064
1.014LysHis: 1.014 ± 0.382
4.308LysIle: 4.308 ± 0.869
5.829LysLys: 5.829 ± 1.694
5.575LysLeu: 5.575 ± 1.328
3.294LysMet: 3.294 ± 1.498
3.294LysAsn: 3.294 ± 0.937
1.521LysPro: 1.521 ± 0.408
0.507LysGln: 0.507 ± 0.297
3.548LysArg: 3.548 ± 0.669
5.575LysSer: 5.575 ± 1.643
2.534LysThr: 2.534 ± 1.005
4.562LysVal: 4.562 ± 0.919
2.027LysTrp: 2.027 ± 0.775
2.788LysTyr: 2.788 ± 0.713
0.0LysXaa: 0.0 ± 0.0
Leu
3.294LeuAla: 3.294 ± 1.278
1.014LeuCys: 1.014 ± 0.538
7.603LeuAsp: 7.603 ± 1.348
3.548LeuGlu: 3.548 ± 0.814
3.548LeuPhe: 3.548 ± 0.517
4.055LeuGly: 4.055 ± 0.844
2.027LeuHis: 2.027 ± 0.62
5.322LeuIle: 5.322 ± 1.174
7.603LeuLys: 7.603 ± 1.066
11.404LeuLeu: 11.404 ± 1.932
4.815LeuMet: 4.815 ± 1.101
4.055LeuAsn: 4.055 ± 0.742
3.294LeuPro: 3.294 ± 1.04
3.548LeuGln: 3.548 ± 1.215
6.589LeuArg: 6.589 ± 2.238
6.842LeuSer: 6.842 ± 1.251
6.336LeuThr: 6.336 ± 1.036
6.082LeuVal: 6.082 ± 1.071
2.027LeuTrp: 2.027 ± 1.235
3.041LeuTyr: 3.041 ± 0.733
0.0LeuXaa: 0.0 ± 0.0
Met
1.774MetAla: 1.774 ± 0.805
0.253MetCys: 0.253 ± 0.148
1.521MetAsp: 1.521 ± 0.496
1.774MetGlu: 1.774 ± 0.878
1.267MetPhe: 1.267 ± 0.967
1.521MetGly: 1.521 ± 0.89
0.507MetHis: 0.507 ± 0.288
4.308MetIle: 4.308 ± 0.615
4.562MetLys: 4.562 ± 1.508
2.534MetLeu: 2.534 ± 0.625
0.507MetMet: 0.507 ± 0.297
0.76MetAsn: 0.76 ± 0.375
1.267MetPro: 1.267 ± 0.541
1.014MetGln: 1.014 ± 0.373
1.267MetArg: 1.267 ± 0.432
3.294MetSer: 3.294 ± 1.282
1.774MetThr: 1.774 ± 0.491
1.774MetVal: 1.774 ± 1.008
0.76MetTrp: 0.76 ± 0.501
1.014MetTyr: 1.014 ± 0.751
0.0MetXaa: 0.0 ± 0.0
Asn
3.294AsnAla: 3.294 ± 1.066
0.507AsnCys: 0.507 ± 0.433
1.521AsnAsp: 1.521 ± 0.795
1.521AsnGlu: 1.521 ± 0.566
1.014AsnPhe: 1.014 ± 0.376
1.267AsnGly: 1.267 ± 0.967
1.014AsnHis: 1.014 ± 0.314
2.788AsnIle: 2.788 ± 0.639
2.281AsnLys: 2.281 ± 0.863
3.548AsnLeu: 3.548 ± 1.16
2.788AsnMet: 2.788 ± 1.157
1.521AsnAsn: 1.521 ± 1.282
1.267AsnPro: 1.267 ± 0.487
1.774AsnGln: 1.774 ± 0.682
1.014AsnArg: 1.014 ± 0.442
5.068AsnSer: 5.068 ± 1.152
3.294AsnThr: 3.294 ± 0.768
2.534AsnVal: 2.534 ± 0.717
0.76AsnTrp: 0.76 ± 0.314
2.281AsnTyr: 2.281 ± 0.782
0.0AsnXaa: 0.0 ± 0.0
Pro
3.294ProAla: 3.294 ± 1.014
0.253ProCys: 0.253 ± 0.148
1.521ProAsp: 1.521 ± 0.546
1.014ProGlu: 1.014 ± 0.577
1.267ProPhe: 1.267 ± 0.608
2.788ProGly: 2.788 ± 0.742
1.774ProHis: 1.774 ± 0.726
2.281ProIle: 2.281 ± 1.05
2.788ProLys: 2.788 ± 0.906
3.041ProLeu: 3.041 ± 1.217
1.267ProMet: 1.267 ± 0.528
1.267ProAsn: 1.267 ± 0.432
1.521ProPro: 1.521 ± 0.89
1.014ProGln: 1.014 ± 0.439
2.027ProArg: 2.027 ± 0.65
4.562ProSer: 4.562 ± 1.155
3.548ProThr: 3.548 ± 0.549
1.014ProVal: 1.014 ± 0.502
0.76ProTrp: 0.76 ± 0.663
2.027ProTyr: 2.027 ± 0.606
0.0ProXaa: 0.0 ± 0.0
Gln
1.267GlnAla: 1.267 ± 0.683
0.507GlnCys: 0.507 ± 0.309
2.281GlnAsp: 2.281 ± 0.768
2.534GlnGlu: 2.534 ± 1.041
0.507GlnPhe: 0.507 ± 0.297
2.027GlnGly: 2.027 ± 0.451
1.267GlnHis: 1.267 ± 0.38
2.281GlnIle: 2.281 ± 0.907
0.76GlnLys: 0.76 ± 0.4
2.788GlnLeu: 2.788 ± 1.494
1.521GlnMet: 1.521 ± 0.683
0.76GlnAsn: 0.76 ± 0.314
0.507GlnPro: 0.507 ± 0.609
0.253GlnGln: 0.253 ± 0.148
1.521GlnArg: 1.521 ± 0.548
1.014GlnSer: 1.014 ± 0.566
1.774GlnThr: 1.774 ± 0.59
1.774GlnVal: 1.774 ± 0.507
0.253GlnTrp: 0.253 ± 0.391
1.521GlnTyr: 1.521 ± 0.725
0.0GlnXaa: 0.0 ± 0.0
Arg
3.548ArgAla: 3.548 ± 0.456
0.76ArgCys: 0.76 ± 0.663
2.281ArgAsp: 2.281 ± 0.951
3.294ArgGlu: 3.294 ± 0.854
2.788ArgPhe: 2.788 ± 0.573
2.788ArgGly: 2.788 ± 0.709
1.014ArgHis: 1.014 ± 0.873
2.281ArgIle: 2.281 ± 0.689
3.294ArgLys: 3.294 ± 0.66
5.575ArgLeu: 5.575 ± 0.854
1.521ArgMet: 1.521 ± 0.66
1.521ArgAsn: 1.521 ± 0.789
2.281ArgPro: 2.281 ± 0.826
1.267ArgGln: 1.267 ± 0.478
2.788ArgArg: 2.788 ± 0.746
5.068ArgSer: 5.068 ± 1.02
4.562ArgThr: 4.562 ± 0.825
3.801ArgVal: 3.801 ± 0.933
0.0ArgTrp: 0.0 ± 0.0
4.055ArgTyr: 4.055 ± 1.011
0.0ArgXaa: 0.0 ± 0.0
Ser
3.294SerAla: 3.294 ± 1.131
2.788SerCys: 2.788 ± 1.62
4.562SerAsp: 4.562 ± 1.653
6.336SerGlu: 6.336 ± 1.231
3.041SerPhe: 3.041 ± 1.348
7.349SerGly: 7.349 ± 1.443
1.774SerHis: 1.774 ± 0.895
4.055SerIle: 4.055 ± 1.031
5.068SerLys: 5.068 ± 0.8
9.123SerLeu: 9.123 ± 1.487
2.788SerMet: 2.788 ± 1.265
4.308SerAsn: 4.308 ± 0.64
3.041SerPro: 3.041 ± 0.867
2.027SerGln: 2.027 ± 0.735
5.322SerArg: 5.322 ± 1.43
8.616SerSer: 8.616 ± 2.282
4.562SerThr: 4.562 ± 1.029
5.829SerVal: 5.829 ± 1.139
1.774SerTrp: 1.774 ± 0.484
2.027SerTyr: 2.027 ± 0.881
0.0SerXaa: 0.0 ± 0.0
Thr
3.801ThrAla: 3.801 ± 0.478
0.507ThrCys: 0.507 ± 0.309
2.281ThrAsp: 2.281 ± 0.67
4.055ThrGlu: 4.055 ± 1.655
3.294ThrPhe: 3.294 ± 0.564
4.308ThrGly: 4.308 ± 2.285
1.521ThrHis: 1.521 ± 0.926
5.322ThrIle: 5.322 ± 1.105
3.041ThrLys: 3.041 ± 1.229
4.308ThrLeu: 4.308 ± 0.843
1.521ThrMet: 1.521 ± 0.699
0.76ThrAsn: 0.76 ± 0.663
1.774ThrPro: 1.774 ± 0.62
2.027ThrGln: 2.027 ± 0.644
2.788ThrArg: 2.788 ± 1.642
5.575ThrSer: 5.575 ± 2.609
1.774ThrThr: 1.774 ± 0.716
4.562ThrVal: 4.562 ± 1.315
1.014ThrTrp: 1.014 ± 0.668
3.041ThrTyr: 3.041 ± 0.928
0.0ThrXaa: 0.0 ± 0.0
Val
3.294ValAla: 3.294 ± 1.275
1.267ValCys: 1.267 ± 0.369
4.055ValAsp: 4.055 ± 1.213
6.842ValGlu: 6.842 ± 2.217
2.788ValPhe: 2.788 ± 0.526
3.041ValGly: 3.041 ± 0.604
0.507ValHis: 0.507 ± 0.361
3.548ValIle: 3.548 ± 0.904
2.788ValLys: 2.788 ± 0.87
6.842ValLeu: 6.842 ± 1.955
1.267ValMet: 1.267 ± 0.58
3.041ValAsn: 3.041 ± 0.709
3.548ValPro: 3.548 ± 0.795
1.014ValGln: 1.014 ± 0.538
3.041ValArg: 3.041 ± 0.969
5.322ValSer: 5.322 ± 1.281
4.562ValThr: 4.562 ± 1.125
3.041ValVal: 3.041 ± 0.837
1.014ValTrp: 1.014 ± 0.382
1.774ValTyr: 1.774 ± 0.696
0.0ValXaa: 0.0 ± 0.0
Trp
1.521TrpAla: 1.521 ± 0.624
0.0TrpCys: 0.0 ± 0.0
0.76TrpAsp: 0.76 ± 0.314
1.774TrpGlu: 1.774 ± 0.595
0.76TrpPhe: 0.76 ± 0.445
0.507TrpGly: 0.507 ± 0.297
0.0TrpHis: 0.0 ± 0.0
1.267TrpIle: 1.267 ± 0.86
1.521TrpLys: 1.521 ± 0.462
1.521TrpLeu: 1.521 ± 0.592
0.507TrpMet: 0.507 ± 0.297
1.014TrpAsn: 1.014 ± 0.733
0.0TrpPro: 0.0 ± 0.0
0.253TrpGln: 0.253 ± 0.148
1.267TrpArg: 1.267 ± 0.606
0.507TrpSer: 0.507 ± 0.309
1.267TrpThr: 1.267 ± 0.608
0.76TrpVal: 0.76 ± 0.4
0.0TrpTrp: 0.0 ± 0.0
0.507TrpTyr: 0.507 ± 0.633
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.281TyrAla: 2.281 ± 0.81
1.014TyrCys: 1.014 ± 0.694
1.521TyrAsp: 1.521 ± 0.842
2.027TyrGlu: 2.027 ± 0.808
1.774TyrPhe: 1.774 ± 0.818
2.281TyrGly: 2.281 ± 0.783
2.027TyrHis: 2.027 ± 0.617
1.521TyrIle: 1.521 ± 0.804
1.774TyrLys: 1.774 ± 0.59
3.294TyrLeu: 3.294 ± 1.102
0.76TyrMet: 0.76 ± 0.43
0.76TyrAsn: 0.76 ± 0.363
2.534TyrPro: 2.534 ± 0.975
1.774TyrGln: 1.774 ± 0.568
2.534TyrArg: 2.534 ± 0.797
3.801TyrSer: 3.801 ± 1.021
2.281TyrThr: 2.281 ± 0.693
1.774TyrVal: 1.774 ± 0.682
0.253TyrTrp: 0.253 ± 0.148
2.788TyrTyr: 2.788 ± 0.816
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 10 proteins (3947 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski