Amino acid dipepetide frequency for Pygoscelis adeliae papillomavirus 1

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
5.072AlaAla: 5.072 ± 1.452
0.39AlaCys: 0.39 ± 0.372
3.121AlaAsp: 3.121 ± 0.827
4.292AlaGlu: 4.292 ± 1.435
1.561AlaPhe: 1.561 ± 1.104
5.072AlaGly: 5.072 ± 1.83
1.171AlaHis: 1.171 ± 0.585
1.951AlaIle: 1.951 ± 0.519
2.341AlaLys: 2.341 ± 1.008
4.292AlaLeu: 4.292 ± 1.699
0.0AlaMet: 0.0 ± 0.0
2.731AlaAsn: 2.731 ± 1.098
8.194AlaPro: 8.194 ± 1.5
1.951AlaGln: 1.951 ± 0.797
3.121AlaArg: 3.121 ± 1.009
3.512AlaSer: 3.512 ± 1.24
3.512AlaThr: 3.512 ± 0.94
4.682AlaVal: 4.682 ± 0.814
0.0AlaTrp: 0.0 ± 0.0
1.951AlaTyr: 1.951 ± 0.559
0.0AlaXaa: 0.0 ± 0.0
Cys
1.561CysAla: 1.561 ± 0.852
0.39CysCys: 0.39 ± 0.422
1.171CysAsp: 1.171 ± 0.675
0.78CysGlu: 0.78 ± 0.59
1.561CysPhe: 1.561 ± 0.636
3.121CysGly: 3.121 ± 0.815
0.39CysHis: 0.39 ± 0.472
1.951CysIle: 1.951 ± 1.015
1.171CysLys: 1.171 ± 0.61
0.78CysLeu: 0.78 ± 0.406
0.0CysMet: 0.0 ± 0.0
1.951CysAsn: 1.951 ± 1.368
1.171CysPro: 1.171 ± 0.719
0.78CysGln: 0.78 ± 0.662
0.78CysArg: 0.78 ± 0.556
1.171CysSer: 1.171 ± 0.585
3.121CysThr: 3.121 ± 1.165
0.0CysVal: 0.0 ± 0.0
0.39CysTrp: 0.39 ± 0.3
1.171CysTyr: 1.171 ± 1.121
0.0CysXaa: 0.0 ± 0.0
Asp
1.951AspAla: 1.951 ± 0.378
2.731AspCys: 2.731 ± 0.982
5.853AspAsp: 5.853 ± 1.473
1.951AspGlu: 1.951 ± 0.597
1.951AspPhe: 1.951 ± 1.085
6.243AspGly: 6.243 ± 1.383
0.78AspHis: 0.78 ± 0.601
5.853AspIle: 5.853 ± 0.59
1.171AspLys: 1.171 ± 0.719
6.633AspLeu: 6.633 ± 1.868
2.341AspMet: 2.341 ± 0.995
3.902AspAsn: 3.902 ± 1.294
6.633AspPro: 6.633 ± 1.843
2.341AspGln: 2.341 ± 1.188
4.292AspArg: 4.292 ± 1.38
3.902AspSer: 3.902 ± 2.436
5.853AspThr: 5.853 ± 0.839
3.512AspVal: 3.512 ± 1.014
0.39AspTrp: 0.39 ± 0.3
2.731AspTyr: 2.731 ± 0.505
0.0AspXaa: 0.0 ± 0.0
Glu
5.853GluAla: 5.853 ± 1.762
1.561GluCys: 1.561 ± 1.045
4.682GluAsp: 4.682 ± 1.752
7.803GluGlu: 7.803 ± 2.679
1.561GluPhe: 1.561 ± 0.637
5.462GluGly: 5.462 ± 1.74
2.341GluHis: 2.341 ± 1.118
1.951GluIle: 1.951 ± 0.784
1.171GluLys: 1.171 ± 0.749
6.633GluLeu: 6.633 ± 1.538
1.561GluMet: 1.561 ± 0.702
2.341GluAsn: 2.341 ± 1.071
0.78GluPro: 0.78 ± 0.68
1.171GluGln: 1.171 ± 0.425
2.731GluArg: 2.731 ± 1.025
2.341GluSer: 2.341 ± 0.951
3.512GluThr: 3.512 ± 1.504
1.951GluVal: 1.951 ± 0.766
0.39GluTrp: 0.39 ± 0.3
1.171GluTyr: 1.171 ± 0.585
0.0GluXaa: 0.0 ± 0.0
Phe
1.171PheAla: 1.171 ± 1.021
0.78PheCys: 0.78 ± 0.783
1.171PheAsp: 1.171 ± 0.535
1.951PheGlu: 1.951 ± 1.286
1.171PhePhe: 1.171 ± 0.61
1.951PheGly: 1.951 ± 0.41
0.0PheHis: 0.0 ± 0.0
1.951PheIle: 1.951 ± 0.89
2.341PheLys: 2.341 ± 1.031
3.512PheLeu: 3.512 ± 1.399
0.0PheMet: 0.0 ± 0.0
1.951PheAsn: 1.951 ± 0.797
1.951PhePro: 1.951 ± 0.928
0.78PheGln: 0.78 ± 0.372
1.171PheArg: 1.171 ± 0.605
1.561PheSer: 1.561 ± 0.581
1.171PheThr: 1.171 ± 0.394
1.951PheVal: 1.951 ± 0.878
0.78PheTrp: 0.78 ± 0.406
1.561PheTyr: 1.561 ± 0.987
0.0PheXaa: 0.0 ± 0.0
Gly
5.853GlyAla: 5.853 ± 1.767
0.78GlyCys: 0.78 ± 0.406
5.853GlyAsp: 5.853 ± 1.254
4.292GlyGlu: 4.292 ± 1.405
1.951GlyPhe: 1.951 ± 0.64
11.315GlyGly: 11.315 ± 2.487
1.561GlyHis: 1.561 ± 0.563
1.951GlyIle: 1.951 ± 1.063
0.78GlyLys: 0.78 ± 0.429
6.243GlyLeu: 6.243 ± 1.599
1.951GlyMet: 1.951 ± 1.045
2.341GlyAsn: 2.341 ± 0.931
6.633GlyPro: 6.633 ± 1.601
3.121GlyGln: 3.121 ± 1.053
4.292GlyArg: 4.292 ± 1.059
2.341GlySer: 2.341 ± 1.626
5.853GlyThr: 5.853 ± 1.942
5.462GlyVal: 5.462 ± 1.576
1.951GlyTrp: 1.951 ± 0.747
1.171GlyTyr: 1.171 ± 0.609
0.0GlyXaa: 0.0 ± 0.0
His
0.78HisAla: 0.78 ± 0.372
1.171HisCys: 1.171 ± 0.719
0.39HisAsp: 0.39 ± 0.34
0.78HisGlu: 0.78 ± 0.601
0.78HisPhe: 0.78 ± 0.372
1.561HisGly: 1.561 ± 0.706
0.39HisHis: 0.39 ± 0.372
1.171HisIle: 1.171 ± 0.61
0.39HisLys: 0.39 ± 0.3
3.512HisLeu: 3.512 ± 0.885
0.78HisMet: 0.78 ± 0.601
0.0HisAsn: 0.0 ± 0.0
0.78HisPro: 0.78 ± 0.42
0.78HisGln: 0.78 ± 0.429
2.341HisArg: 2.341 ± 0.715
0.78HisSer: 0.78 ± 0.662
3.121HisThr: 3.121 ± 1.221
0.78HisVal: 0.78 ± 0.523
0.78HisTrp: 0.78 ± 0.473
0.78HisTyr: 0.78 ± 0.601
0.0HisXaa: 0.0 ± 0.0
Ile
2.341IleAla: 2.341 ± 1.047
1.561IleCys: 1.561 ± 0.6
3.121IleAsp: 3.121 ± 0.733
2.731IleGlu: 2.731 ± 0.945
2.341IlePhe: 2.341 ± 0.841
2.341IleGly: 2.341 ± 0.627
0.39IleHis: 0.39 ± 0.34
1.561IleIle: 1.561 ± 0.905
0.0IleLys: 0.0 ± 0.0
2.341IleLeu: 2.341 ± 0.857
0.39IleMet: 0.39 ± 0.372
1.171IleAsn: 1.171 ± 0.425
5.072IlePro: 5.072 ± 1.802
1.561IleGln: 1.561 ± 0.64
1.561IleArg: 1.561 ± 0.501
4.292IleSer: 4.292 ± 1.564
3.121IleThr: 3.121 ± 0.808
1.561IleVal: 1.561 ± 0.581
0.0IleTrp: 0.0 ± 0.0
1.171IleTyr: 1.171 ± 0.584
0.0IleXaa: 0.0 ± 0.0
Lys
1.951LysAla: 1.951 ± 0.768
1.171LysCys: 1.171 ± 0.585
2.341LysAsp: 2.341 ± 1.233
1.171LysGlu: 1.171 ± 0.778
0.0LysPhe: 0.0 ± 0.0
1.561LysGly: 1.561 ± 0.695
0.39LysHis: 0.39 ± 0.372
0.0LysIle: 0.0 ± 0.0
1.171LysLys: 1.171 ± 0.778
2.731LysLeu: 2.731 ± 1.36
0.0LysMet: 0.0 ± 0.0
1.171LysAsn: 1.171 ± 0.61
1.171LysPro: 1.171 ± 0.62
0.39LysGln: 0.39 ± 0.372
5.072LysArg: 5.072 ± 1.445
1.951LysSer: 1.951 ± 1.152
1.951LysThr: 1.951 ± 0.726
1.171LysVal: 1.171 ± 1.117
0.39LysTrp: 0.39 ± 0.404
3.512LysTyr: 3.512 ± 1.447
0.0LysXaa: 0.0 ± 0.0
Leu
2.731LeuAla: 2.731 ± 0.696
2.731LeuCys: 2.731 ± 1.176
5.072LeuAsp: 5.072 ± 0.549
5.072LeuGlu: 5.072 ± 0.996
3.121LeuPhe: 3.121 ± 0.543
5.462LeuGly: 5.462 ± 1.406
1.171LeuHis: 1.171 ± 0.668
3.902LeuIle: 3.902 ± 1.158
3.512LeuLys: 3.512 ± 1.336
8.584LeuLeu: 8.584 ± 2.015
1.951LeuMet: 1.951 ± 0.818
3.121LeuAsn: 3.121 ± 0.947
8.194LeuPro: 8.194 ± 2.699
6.243LeuGln: 6.243 ± 1.602
5.462LeuArg: 5.462 ± 2.207
8.194LeuSer: 8.194 ± 1.785
7.413LeuThr: 7.413 ± 1.193
3.512LeuVal: 3.512 ± 1.19
3.121LeuTrp: 3.121 ± 0.639
5.072LeuTyr: 5.072 ± 1.137
0.0LeuXaa: 0.0 ± 0.0
Met
0.78MetAla: 0.78 ± 0.45
1.171MetCys: 1.171 ± 1.243
1.561MetAsp: 1.561 ± 0.442
1.171MetGlu: 1.171 ± 0.719
0.78MetPhe: 0.78 ± 0.372
0.78MetGly: 0.78 ± 0.45
0.78MetHis: 0.78 ± 0.473
0.39MetIle: 0.39 ± 0.3
0.0MetLys: 0.0 ± 0.0
1.171MetLeu: 1.171 ± 0.535
0.0MetMet: 0.0 ± 0.0
0.39MetAsn: 0.39 ± 0.3
1.171MetPro: 1.171 ± 0.64
1.171MetGln: 1.171 ± 0.647
1.951MetArg: 1.951 ± 0.929
2.731MetSer: 2.731 ± 1.428
0.78MetThr: 0.78 ± 0.574
1.561MetVal: 1.561 ± 0.696
0.0MetTrp: 0.0 ± 0.0
0.78MetTyr: 0.78 ± 0.473
0.0MetXaa: 0.0 ± 0.0
Asn
1.561AsnAla: 1.561 ± 0.696
0.39AsnCys: 0.39 ± 0.372
1.951AsnAsp: 1.951 ± 0.986
1.171AsnGlu: 1.171 ± 0.675
0.78AsnPhe: 0.78 ± 0.601
2.341AsnGly: 2.341 ± 0.676
1.171AsnHis: 1.171 ± 0.901
1.561AsnIle: 1.561 ± 0.744
1.171AsnLys: 1.171 ± 0.901
2.731AsnLeu: 2.731 ± 1.307
0.0AsnMet: 0.0 ± 0.0
2.341AsnAsn: 2.341 ± 1.113
3.512AsnPro: 3.512 ± 0.861
1.171AsnGln: 1.171 ± 0.494
2.731AsnArg: 2.731 ± 1.058
3.512AsnSer: 3.512 ± 1.124
5.072AsnThr: 5.072 ± 1.024
1.951AsnVal: 1.951 ± 0.731
0.39AsnTrp: 0.39 ± 0.422
1.171AsnTyr: 1.171 ± 0.61
0.0AsnXaa: 0.0 ± 0.0
Pro
5.072ProAla: 5.072 ± 1.149
0.39ProCys: 0.39 ± 0.3
5.462ProAsp: 5.462 ± 1.298
3.121ProGlu: 3.121 ± 0.929
3.512ProPhe: 3.512 ± 2.13
4.292ProGly: 4.292 ± 1.239
1.171ProHis: 1.171 ± 0.82
1.171ProIle: 1.171 ± 0.495
3.512ProLys: 3.512 ± 1.531
8.584ProLeu: 8.584 ± 2.213
2.341ProMet: 2.341 ± 0.588
3.121ProAsn: 3.121 ± 0.815
8.194ProPro: 8.194 ± 3.054
3.512ProGln: 3.512 ± 0.781
5.072ProArg: 5.072 ± 0.844
7.023ProSer: 7.023 ± 1.966
5.853ProThr: 5.853 ± 1.759
3.902ProVal: 3.902 ± 1.46
1.561ProTrp: 1.561 ± 0.718
3.902ProTyr: 3.902 ± 1.038
0.0ProXaa: 0.0 ± 0.0
Gln
2.731GlnAla: 2.731 ± 0.952
0.0GlnCys: 0.0 ± 0.0
2.341GlnAsp: 2.341 ± 0.427
3.512GlnGlu: 3.512 ± 1.723
0.78GlnPhe: 0.78 ± 0.406
1.171GlnGly: 1.171 ± 0.429
1.951GlnHis: 1.951 ± 0.647
0.78GlnIle: 0.78 ± 0.406
1.171GlnLys: 1.171 ± 0.778
5.853GlnLeu: 5.853 ± 0.988
0.39GlnMet: 0.39 ± 0.372
0.78GlnAsn: 0.78 ± 0.745
3.121GlnPro: 3.121 ± 0.584
1.951GlnGln: 1.951 ± 0.929
3.512GlnArg: 3.512 ± 1.518
3.121GlnSer: 3.121 ± 0.968
2.731GlnThr: 2.731 ± 1.305
1.561GlnVal: 1.561 ± 0.807
1.171GlnTrp: 1.171 ± 0.634
0.78GlnTyr: 0.78 ± 0.406
0.0GlnXaa: 0.0 ± 0.0
Arg
5.462ArgAla: 5.462 ± 1.181
1.951ArgCys: 1.951 ± 0.939
4.292ArgAsp: 4.292 ± 1.317
3.512ArgGlu: 3.512 ± 1.114
1.171ArgPhe: 1.171 ± 0.425
5.462ArgGly: 5.462 ± 1.938
1.951ArgHis: 1.951 ± 0.647
2.731ArgIle: 2.731 ± 1.018
3.902ArgLys: 3.902 ± 0.777
8.974ArgLeu: 8.974 ± 1.897
3.512ArgMet: 3.512 ± 1.22
1.951ArgAsn: 1.951 ± 0.811
6.243ArgPro: 6.243 ± 1.452
1.951ArgGln: 1.951 ± 0.996
9.364ArgArg: 9.364 ± 2.332
5.853ArgSer: 5.853 ± 1.308
2.731ArgThr: 2.731 ± 1.134
3.121ArgVal: 3.121 ± 0.657
0.78ArgTrp: 0.78 ± 0.372
2.341ArgTyr: 2.341 ± 0.8
0.0ArgXaa: 0.0 ± 0.0
Ser
3.902SerAla: 3.902 ± 1.176
1.171SerCys: 1.171 ± 0.798
6.633SerAsp: 6.633 ± 2.834
2.731SerGlu: 2.731 ± 0.976
1.171SerPhe: 1.171 ± 0.585
5.853SerGly: 5.853 ± 1.277
2.341SerHis: 2.341 ± 0.627
1.951SerIle: 1.951 ± 1.11
2.731SerLys: 2.731 ± 0.591
4.682SerLeu: 4.682 ± 1.521
1.171SerMet: 1.171 ± 0.368
1.561SerAsn: 1.561 ± 0.634
6.243SerPro: 6.243 ± 2.121
2.341SerGln: 2.341 ± 1.439
5.072SerArg: 5.072 ± 0.948
4.292SerSer: 4.292 ± 2.097
6.633SerThr: 6.633 ± 1.513
3.902SerVal: 3.902 ± 0.735
0.39SerTrp: 0.39 ± 0.422
2.341SerTyr: 2.341 ± 0.632
0.0SerXaa: 0.0 ± 0.0
Thr
4.682ThrAla: 4.682 ± 1.452
1.171ThrCys: 1.171 ± 0.72
7.803ThrAsp: 7.803 ± 1.675
6.243ThrGlu: 6.243 ± 1.914
1.951ThrPhe: 1.951 ± 1.213
4.682ThrGly: 4.682 ± 0.966
1.171ThrHis: 1.171 ± 0.647
3.902ThrIle: 3.902 ± 0.4
0.0ThrLys: 0.0 ± 0.0
6.243ThrLeu: 6.243 ± 2.003
1.561ThrMet: 1.561 ± 0.858
3.121ThrAsn: 3.121 ± 1.055
4.292ThrPro: 4.292 ± 0.573
3.512ThrGln: 3.512 ± 1.288
7.413ThrArg: 7.413 ± 2.07
4.682ThrSer: 4.682 ± 0.702
3.121ThrThr: 3.121 ± 0.807
6.243ThrVal: 6.243 ± 0.99
1.561ThrTrp: 1.561 ± 0.706
2.341ThrTyr: 2.341 ± 0.676
0.0ThrXaa: 0.0 ± 0.0
Val
3.121ValAla: 3.121 ± 0.637
1.171ValCys: 1.171 ± 0.62
3.121ValAsp: 3.121 ± 0.966
2.731ValGlu: 2.731 ± 1.106
1.171ValPhe: 1.171 ± 0.429
4.292ValGly: 4.292 ± 0.944
1.171ValHis: 1.171 ± 0.538
3.121ValIle: 3.121 ± 0.815
0.78ValLys: 0.78 ± 0.45
4.682ValLeu: 4.682 ± 1.025
0.39ValMet: 0.39 ± 0.372
1.171ValAsn: 1.171 ± 0.717
4.682ValPro: 4.682 ± 1.62
1.951ValGln: 1.951 ± 0.64
5.462ValArg: 5.462 ± 1.332
2.731ValSer: 2.731 ± 0.673
6.243ValThr: 6.243 ± 1.308
1.561ValVal: 1.561 ± 0.271
0.39ValTrp: 0.39 ± 0.372
1.561ValTyr: 1.561 ± 0.732
0.0ValXaa: 0.0 ± 0.0
Trp
0.78TrpAla: 0.78 ± 0.372
0.39TrpCys: 0.39 ± 0.3
1.561TrpAsp: 1.561 ± 0.744
0.78TrpGlu: 0.78 ± 0.509
0.0TrpPhe: 0.0 ± 0.0
0.78TrpGly: 0.78 ± 0.473
0.78TrpHis: 0.78 ± 0.645
0.0TrpIle: 0.0 ± 0.0
0.78TrpLys: 0.78 ± 0.429
2.341TrpLeu: 2.341 ± 0.75
0.0TrpMet: 0.0 ± 0.0
0.39TrpAsn: 0.39 ± 0.422
0.78TrpPro: 0.78 ± 0.372
1.171TrpGln: 1.171 ± 0.749
1.951TrpArg: 1.951 ± 0.782
0.39TrpSer: 0.39 ± 0.372
1.951TrpThr: 1.951 ± 1.264
0.78TrpVal: 0.78 ± 0.601
0.0TrpTrp: 0.0 ± 0.0
0.39TrpTyr: 0.39 ± 0.404
0.0TrpXaa: 0.0 ± 0.0
Tyr
1.561TyrAla: 1.561 ± 0.873
2.341TyrCys: 2.341 ± 0.85
3.512TyrAsp: 3.512 ± 0.607
1.171TyrGlu: 1.171 ± 0.668
1.561TyrPhe: 1.561 ± 0.553
1.951TyrGly: 1.951 ± 0.526
0.78TyrHis: 0.78 ± 0.406
0.78TyrIle: 0.78 ± 0.42
1.171TyrLys: 1.171 ± 0.901
3.121TyrLeu: 3.121 ± 0.742
0.39TyrMet: 0.39 ± 0.491
1.171TyrAsn: 1.171 ± 0.64
1.951TyrPro: 1.951 ± 0.996
1.561TyrGln: 1.561 ± 0.982
3.902TyrArg: 3.902 ± 1.09
2.731TyrSer: 2.731 ± 0.923
1.951TyrThr: 1.951 ± 1.11
2.341TyrVal: 2.341 ± 1.004
1.561TyrTrp: 1.561 ± 0.723
2.731TyrTyr: 2.731 ± 0.994
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (2564 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski