Amino acid dipepetide frequency for Beauveria bassiana (strain ARSEF 2860) (White muscardine disease fungus) (Tritirachium shiotae)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10^-3.

For more information see sequence space article on Wikipedia.

	Ala	Cys	Asp	Glu	Phe	Gly	His	Ile	Lys	Leu	Met	Asn	Pro	Gln	Arg	Ser	Thr	Val	Trp	Tyr	Xaa
Ala	12.266AlaAla: 12.266 ± 0.079	1.205AlaCys: 1.205 ± 0.016	5.029AlaAsp: 5.029 ± 0.033	5.499AlaGlu: 5.499 ± 0.038	3.247AlaPhe: 3.247 ± 0.028	6.436AlaGly: 6.436 ± 0.041	2.033AlaHis: 2.033 ± 0.017	4.342AlaIle: 4.342 ± 0.031	4.39AlaLys: 4.39 ± 0.034	8.421AlaLeu: 8.421 ± 0.046	2.207AlaMet: 2.207 ± 0.024	3.084AlaAsn: 3.084 ± 0.027	5.17AlaPro: 5.17 ± 0.043	3.733AlaGln: 3.733 ± 0.029	5.556AlaArg: 5.556 ± 0.039	7.918AlaSer: 7.918 ± 0.048	5.993AlaThr: 5.993 ± 0.038	6.174AlaVal: 6.174 ± 0.045	1.276AlaTrp: 1.276 ± 0.018	2.221AlaTyr: 2.221 ± 0.022	0.0AlaXaa: 0.0 ± 0.0
Cys	1.1CysAla: 1.1 ± 0.018	0.252CysCys: 0.252 ± 0.007	0.711CysAsp: 0.711 ± 0.012	0.622CysGlu: 0.622 ± 0.012	0.552CysPhe: 0.552 ± 0.01	0.984CysGly: 0.984 ± 0.016	0.339CysHis: 0.339 ± 0.009	0.707CysIle: 0.707 ± 0.013	0.553CysLys: 0.553 ± 0.011	1.307CysLeu: 1.307 ± 0.017	0.277CysMet: 0.277 ± 0.007	0.463CysAsn: 0.463 ± 0.012	0.691CysPro: 0.691 ± 0.015	0.475CysGln: 0.475 ± 0.009	0.831CysArg: 0.831 ± 0.013	0.927CysSer: 0.927 ± 0.014	0.711CysThr: 0.711 ± 0.013	0.826CysVal: 0.826 ± 0.014	0.204CysTrp: 0.204 ± 0.007	0.349CysTyr: 0.349 ± 0.008	0.0CysXaa: 0.0 ± 0.0
Asp	5.72AspAla: 5.72 ± 0.039	0.717AspCys: 0.717 ± 0.014	4.878AspAsp: 4.878 ± 0.053	4.637AspGlu: 4.637 ± 0.038	2.202AspPhe: 2.202 ± 0.022	4.514AspGly: 4.514 ± 0.032	1.212AspHis: 1.212 ± 0.017	2.863AspIle: 2.863 ± 0.025	2.642AspLys: 2.642 ± 0.027	4.81AspLeu: 4.81 ± 0.034	1.422AspMet: 1.422 ± 0.017	1.958AspAsn: 1.958 ± 0.022	2.967AspPro: 2.967 ± 0.024	1.802AspGln: 1.802 ± 0.017	3.082AspArg: 3.082 ± 0.037	4.117AspSer: 4.117 ± 0.035	2.977AspThr: 2.977 ± 0.026	3.827AspVal: 3.827 ± 0.025	0.899AspTrp: 0.899 ± 0.014	1.583AspTyr: 1.583 ± 0.019	0.0AspXaa: 0.0 ± 0.0
Glu	5.76GluAla: 5.76 ± 0.044	0.641GluCys: 0.641 ± 0.012	3.943GluAsp: 3.943 ± 0.038	4.963GluGlu: 4.963 ± 0.059	1.879GluPhe: 1.879 ± 0.02	3.263GluGly: 3.263 ± 0.03	1.407GluHis: 1.407 ± 0.016	2.706GluIle: 2.706 ± 0.024	3.3GluLys: 3.3 ± 0.033	5.202GluLeu: 5.202 ± 0.038	1.464GluMet: 1.464 ± 0.019	2.103GluAsn: 2.103 ± 0.023	2.784GluPro: 2.784 ± 0.032	2.576GluGln: 2.576 ± 0.025	3.758GluArg: 3.758 ± 0.035	4.046GluSer: 4.046 ± 0.036	3.462GluThr: 3.462 ± 0.03	3.202GluVal: 3.202 ± 0.024	0.835GluTrp: 0.835 ± 0.013	1.58GluTyr: 1.58 ± 0.017	0.0GluXaa: 0.0 ± 0.0
Phe	3.309PheAla: 3.309 ± 0.028	0.583PheCys: 0.583 ± 0.011	2.348PheAsp: 2.348 ± 0.023	2.026PheGlu: 2.026 ± 0.019	1.58PhePhe: 1.58 ± 0.02	2.818PheGly: 2.818 ± 0.026	0.87PheHis: 0.87 ± 0.014	1.689PheIle: 1.689 ± 0.021	1.432PheLys: 1.432 ± 0.017	3.337PheLeu: 3.337 ± 0.03	0.781PheMet: 0.781 ± 0.013	1.327PheAsn: 1.327 ± 0.015	1.778PhePro: 1.778 ± 0.018	1.348PheGln: 1.348 ± 0.016	1.957PheArg: 1.957 ± 0.021	2.857PheSer: 2.857 ± 0.024	2.073PheThr: 2.073 ± 0.021	2.377PheVal: 2.377 ± 0.024	0.631PheTrp: 0.631 ± 0.012	1.075PheTyr: 1.075 ± 0.014	0.0PheXaa: 0.0 ± 0.0
Gly	5.932GlyAla: 5.932 ± 0.044	0.88GlyCys: 0.88 ± 0.016	3.83GlyAsp: 3.83 ± 0.03	3.5GlyGlu: 3.5 ± 0.026	2.7GlyPhe: 2.7 ± 0.024	6.319GlyGly: 6.319 ± 0.063	1.749GlyHis: 1.749 ± 0.021	3.424GlyIle: 3.424 ± 0.032	3.407GlyLys: 3.407 ± 0.032	5.949GlyLeu: 5.949 ± 0.038	1.561GlyMet: 1.561 ± 0.019	2.549GlyAsn: 2.549 ± 0.029	3.228GlyPro: 3.228 ± 0.03	2.636GlyGln: 2.636 ± 0.025	4.343GlyArg: 4.343 ± 0.035	5.728GlySer: 5.728 ± 0.041	3.837GlyThr: 3.837 ± 0.031	4.274GlyVal: 4.274 ± 0.03	1.151GlyTrp: 1.151 ± 0.016	2.077GlyTyr: 2.077 ± 0.025	0.0GlyXaa: 0.0 ± 0.0
His	2.084HisAla: 2.084 ± 0.024	0.359HisCys: 0.359 ± 0.009	1.428HisAsp: 1.428 ± 0.016	1.317HisGlu: 1.317 ± 0.018	0.92HisPhe: 0.92 ± 0.015	1.859HisGly: 1.859 ± 0.021	0.939HisHis: 0.939 ± 0.022	1.19HisIle: 1.19 ± 0.014	0.923HisLys: 0.923 ± 0.013	2.247HisLeu: 2.247 ± 0.02	0.551HisMet: 0.551 ± 0.011	0.867HisAsn: 0.867 ± 0.015	1.508HisPro: 1.508 ± 0.021	1.034HisGln: 1.034 ± 0.018	1.623HisArg: 1.623 ± 0.021	1.776HisSer: 1.776 ± 0.025	1.231HisThr: 1.231 ± 0.015	1.517HisVal: 1.517 ± 0.018	0.351HisTrp: 0.351 ± 0.009	0.675HisTyr: 0.675 ± 0.011	0.0HisXaa: 0.0 ± 0.0
Ile	4.259IleAla: 4.259 ± 0.036	0.749IleCys: 0.749 ± 0.012	2.802IleAsp: 2.802 ± 0.024	2.655IleGlu: 2.655 ± 0.026	1.856IlePhe: 1.856 ± 0.021	3.126IleGly: 3.126 ± 0.025	1.147IleHis: 1.147 ± 0.016	2.338IleIle: 2.338 ± 0.023	2.126IleLys: 2.126 ± 0.024	4.291IleLeu: 4.291 ± 0.035	1.038IleMet: 1.038 ± 0.015	1.671IleAsn: 1.671 ± 0.02	2.663IlePro: 2.663 ± 0.024	1.751IleGln: 1.751 ± 0.021	2.707IleArg: 2.707 ± 0.022	3.495IleSer: 3.495 ± 0.029	2.666IleThr: 2.666 ± 0.025	3.121IleVal: 3.121 ± 0.029	0.676IleTrp: 0.676 ± 0.013	1.31IleTyr: 1.31 ± 0.016	0.0IleXaa: 0.0 ± 0.0
Lys	4.298LysAla: 4.298 ± 0.04	0.506LysCys: 0.506 ± 0.011	2.694LysAsp: 2.694 ± 0.028	3.031LysGlu: 3.031 ± 0.03	1.418LysPhe: 1.418 ± 0.018	2.75LysGly: 2.75 ± 0.026	1.124LysHis: 1.124 ± 0.014	2.112LysIle: 2.112 ± 0.023	3.186LysLys: 3.186 ± 0.045	4.164LysLeu: 4.164 ± 0.031	1.077LysMet: 1.077 ± 0.016	1.684LysAsn: 1.684 ± 0.02	2.638LysPro: 2.638 ± 0.028	1.934LysGln: 1.934 ± 0.022	3.312LysArg: 3.312 ± 0.029	3.423LysSer: 3.423 ± 0.026	2.917LysThr: 2.917 ± 0.029	2.673LysVal: 2.673 ± 0.026	0.644LysTrp: 0.644 ± 0.011	1.31LysTyr: 1.31 ± 0.014	0.0LysXaa: 0.0 ± 0.0
Leu	8.656LeuAla: 8.656 ± 0.046	1.28LeuCys: 1.28 ± 0.017	5.296LeuAsp: 5.296 ± 0.037	5.18LeuGlu: 5.18 ± 0.035	3.317LeuPhe: 3.317 ± 0.03	5.892LeuGly: 5.892 ± 0.037	2.297LeuHis: 2.297 ± 0.019	3.695LeuIle: 3.695 ± 0.031	3.871LeuLys: 3.871 ± 0.035	8.473LeuLeu: 8.473 ± 0.06	1.758LeuMet: 1.758 ± 0.021	2.881LeuAsn: 2.881 ± 0.024	5.453LeuPro: 5.453 ± 0.04	3.956LeuGln: 3.956 ± 0.033	6.028LeuArg: 6.028 ± 0.041	7.026LeuSer: 7.026 ± 0.043	4.705LeuThr: 4.705 ± 0.035	5.606LeuVal: 5.606 ± 0.036	1.22LeuTrp: 1.22 ± 0.017	2.298LeuTyr: 2.298 ± 0.022	0.0LeuXaa: 0.0 ± 0.0
Met	2.453MetAla: 2.453 ± 0.021	0.269MetCys: 0.269 ± 0.008	1.297MetAsp: 1.297 ± 0.016	1.235MetGlu: 1.235 ± 0.016	0.712MetPhe: 0.712 ± 0.012	1.443MetGly: 1.443 ± 0.02	0.539MetHis: 0.539 ± 0.011	0.946MetIle: 0.946 ± 0.015	0.973MetLys: 0.973 ± 0.013	1.979MetLeu: 1.979 ± 0.019	0.627MetMet: 0.627 ± 0.012	0.751MetAsn: 0.751 ± 0.012	1.404MetPro: 1.404 ± 0.017	0.947MetGln: 0.947 ± 0.016	1.386MetArg: 1.386 ± 0.017	1.855MetSer: 1.855 ± 0.019	1.399MetThr: 1.399 ± 0.017	1.31MetVal: 1.31 ± 0.016	0.26MetTrp: 0.26 ± 0.007	0.551MetTyr: 0.551 ± 0.01	0.0MetXaa: 0.0 ± 0.0
Asn	3.178AsnAla: 3.178 ± 0.025	0.451AsnCys: 0.451 ± 0.01	1.989AsnAsp: 1.989 ± 0.021	1.879AsnGlu: 1.879 ± 0.02	1.289AsnPhe: 1.289 ± 0.016	2.941AsnGly: 2.941 ± 0.031	0.845AsnHis: 0.845 ± 0.012	1.884AsnIle: 1.884 ± 0.022	1.593AsnLys: 1.593 ± 0.019	3.044AsnLeu: 3.044 ± 0.026	0.883AsnMet: 0.883 ± 0.012	1.501AsnAsn: 1.501 ± 0.027	2.119AsnPro: 2.119 ± 0.024	1.223AsnGln: 1.223 ± 0.018	1.88AsnArg: 1.88 ± 0.021	2.573AsnSer: 2.573 ± 0.024	2.018AsnThr: 2.018 ± 0.023	2.277AsnVal: 2.277 ± 0.021	0.531AsnTrp: 0.531 ± 0.012	0.985AsnTyr: 0.985 ± 0.013	0.0AsnXaa: 0.0 ± 0.0
Pro	5.737ProAla: 5.737 ± 0.05	0.567ProCys: 0.567 ± 0.01	3.237ProAsp: 3.237 ± 0.029	3.51ProGlu: 3.51 ± 0.029	1.938ProPhe: 1.938 ± 0.019	3.85ProGly: 3.85 ± 0.035	1.298ProHis: 1.298 ± 0.018	2.32ProIle: 2.32 ± 0.025	2.508ProLys: 2.508 ± 0.027	4.637ProLeu: 4.637 ± 0.031	1.085ProMet: 1.085 ± 0.016	1.988ProAsn: 1.988 ± 0.022	5.051ProPro: 5.051 ± 0.078	2.386ProGln: 2.386 ± 0.032	3.483ProArg: 3.483 ± 0.033	5.481ProSer: 5.481 ± 0.049	3.672ProThr: 3.672 ± 0.033	3.541ProVal: 3.541 ± 0.028	0.721ProTrp: 0.721 ± 0.013	1.362ProTyr: 1.362 ± 0.017	0.0ProXaa: 0.0 ± 0.0
Gln	3.612GlnAla: 3.612 ± 0.031	0.461GlnCys: 0.461 ± 0.01	2.214GlnAsp: 2.214 ± 0.02	2.297GlnGlu: 2.297 ± 0.025	1.276GlnPhe: 1.276 ± 0.016	2.408GlnGly: 2.408 ± 0.025	1.211GlnHis: 1.211 ± 0.02	1.791GlnIle: 1.791 ± 0.019	1.904GlnLys: 1.904 ± 0.021	3.636GlnLeu: 3.636 ± 0.033	0.924GlnMet: 0.924 ± 0.015	1.446GlnAsn: 1.446 ± 0.017	2.554GlnPro: 2.554 ± 0.03	2.913GlnGln: 2.913 ± 0.059	2.846GlnArg: 2.846 ± 0.024	3.077GlnSer: 3.077 ± 0.027	2.298GlnThr: 2.298 ± 0.023	2.259GlnVal: 2.259 ± 0.023	0.556GlnTrp: 0.556 ± 0.011	1.133GlnTyr: 1.133 ± 0.017	0.0GlnXaa: 0.0 ± 0.0
Arg	5.31ArgAla: 5.31 ± 0.041	0.808ArgCys: 0.808 ± 0.013	3.663ArgAsp: 3.663 ± 0.034	3.629ArgGlu: 3.629 ± 0.034	2.19ArgPhe: 2.19 ± 0.02	3.988ArgGly: 3.988 ± 0.038	1.716ArgHis: 1.716 ± 0.021	2.895ArgIle: 2.895 ± 0.025	3.234ArgLys: 3.234 ± 0.028	5.721ArgLeu: 5.721 ± 0.037	1.327ArgMet: 1.327 ± 0.016	2.228ArgAsn: 2.228 ± 0.024	3.576ArgPro: 3.576 ± 0.035	2.842ArgGln: 2.842 ± 0.028	5.567ArgArg: 5.567 ± 0.054	4.731ArgSer: 4.731 ± 0.04	3.238ArgThr: 3.238 ± 0.028	3.474ArgVal: 3.474 ± 0.031	0.945ArgTrp: 0.945 ± 0.013	1.632ArgTyr: 1.632 ± 0.019	0.0ArgXaa: 0.0 ± 0.0
Ser	7.221SerAla: 7.221 ± 0.046	0.905SerCys: 0.905 ± 0.017	4.224SerAsp: 4.224 ± 0.035	3.847SerGlu: 3.847 ± 0.033	2.898SerPhe: 2.898 ± 0.026	5.499SerGly: 5.499 ± 0.041	1.913SerHis: 1.913 ± 0.018	3.779SerIle: 3.779 ± 0.031	3.636SerLys: 3.636 ± 0.03	6.978SerLeu: 6.978 ± 0.04	1.714SerMet: 1.714 ± 0.022	2.832SerAsn: 2.832 ± 0.026	5.072SerPro: 5.072 ± 0.05	3.164SerGln: 3.164 ± 0.029	5.045SerArg: 5.045 ± 0.044	8.72SerSer: 8.72 ± 0.08	5.266SerThr: 5.266 ± 0.044	4.603SerVal: 4.603 ± 0.032	1.087SerTrp: 1.087 ± 0.017	1.948SerTyr: 1.948 ± 0.022	0.0SerXaa: 0.0 ± 0.0
Thr	5.805ThrAla: 5.805 ± 0.039	0.749ThrCys: 0.749 ± 0.013	2.889ThrAsp: 2.889 ± 0.027	3.087ThrGlu: 3.087 ± 0.028	2.135ThrPhe: 2.135 ± 0.02	4.053ThrGly: 4.053 ± 0.032	1.253ThrHis: 1.253 ± 0.017	2.913ThrIle: 2.913 ± 0.026	2.56ThrLys: 2.56 ± 0.025	5.146ThrLeu: 5.146 ± 0.032	1.27ThrMet: 1.27 ± 0.016	1.969ThrAsn: 1.969 ± 0.021	4.129ThrPro: 4.129 ± 0.039	2.046ThrGln: 2.046 ± 0.021	3.227ThrArg: 3.227 ± 0.027	5.066ThrSer: 5.066 ± 0.042	4.482ThrThr: 4.482 ± 0.042	3.804ThrVal: 3.804 ± 0.03	0.821ThrTrp: 0.821 ± 0.012	1.481ThrTyr: 1.481 ± 0.018	0.0ThrXaa: 0.0 ± 0.0
Val	6.047ValAla: 6.047 ± 0.04	0.883ValCys: 0.883 ± 0.013	3.828ValAsp: 3.828 ± 0.027	3.732ValGlu: 3.732 ± 0.027	2.431ValPhe: 2.431 ± 0.023	4.038ValGly: 4.038 ± 0.037	1.397ValHis: 1.397 ± 0.016	2.842ValIle: 2.842 ± 0.025	2.77ValLys: 2.77 ± 0.026	5.676ValLeu: 5.676 ± 0.039	1.373ValMet: 1.373 ± 0.016	2.113ValAsn: 2.113 ± 0.021	3.607ValPro: 3.607 ± 0.028	2.408ValGln: 2.408 ± 0.025	3.526ValArg: 3.526 ± 0.026	4.575ValSer: 4.575 ± 0.032	3.499ValThr: 3.499 ± 0.03	4.456ValVal: 4.456 ± 0.034	0.876ValTrp: 0.876 ± 0.014	1.696ValTyr: 1.696 ± 0.02	0.0ValXaa: 0.0 ± 0.0
Trp	1.207TrpAla: 1.207 ± 0.018	0.198TrpCys: 0.198 ± 0.007	0.879TrpAsp: 0.879 ± 0.015	0.79TrpGlu: 0.79 ± 0.013	0.529TrpPhe: 0.529 ± 0.011	0.893TrpGly: 0.893 ± 0.015	0.375TrpHis: 0.375 ± 0.009	0.709TrpIle: 0.709 ± 0.013	0.746TrpLys: 0.746 ± 0.013	1.387TrpLeu: 1.387 ± 0.019	0.353TrpMet: 0.353 ± 0.008	0.589TrpAsn: 0.589 ± 0.012	0.641TrpPro: 0.641 ± 0.012	0.603TrpGln: 0.603 ± 0.011	0.992TrpArg: 0.992 ± 0.015	1.049TrpSer: 1.049 ± 0.014	0.888TrpThr: 0.888 ± 0.014	0.862TrpVal: 0.862 ± 0.014	0.274TrpTrp: 0.274 ± 0.008	0.426TrpTyr: 0.426 ± 0.01	0.0TrpXaa: 0.0 ± 0.0
Tyr	2.249TyrAla: 2.249 ± 0.019	0.418TyrCys: 0.418 ± 0.009	1.65TyrAsp: 1.65 ± 0.018	1.439TyrGlu: 1.439 ± 0.016	1.128TyrPhe: 1.128 ± 0.016	2.0TyrGly: 2.0 ± 0.024	0.735TyrHis: 0.735 ± 0.012	1.302TyrIle: 1.302 ± 0.017	1.14TyrLys: 1.14 ± 0.014	2.482TyrLeu: 2.482 ± 0.021	0.644TyrMet: 0.644 ± 0.013	1.079TyrAsn: 1.079 ± 0.015	1.339TyrPro: 1.339 ± 0.018	1.039TyrGln: 1.039 ± 0.013	1.565TyrArg: 1.565 ± 0.019	1.908TyrSer: 1.908 ± 0.021	1.55TyrThr: 1.55 ± 0.02	1.592TyrVal: 1.592 ± 0.018	0.431TyrTrp: 0.431 ± 0.009	0.906TyrTyr: 0.906 ± 0.015	0.0TyrXaa: 0.0 ± 0.0
Xaa	0.0XaaAla: 0.0 ± 0.0	0.0XaaCys: 0.0 ± 0.0	0.0XaaAsp: 0.0 ± 0.0	0.0XaaGlu: 0.0 ± 0.0	0.0XaaPhe: 0.0 ± 0.0	0.0XaaGly: 0.0 ± 0.0	0.0XaaHis: 0.0 ± 0.0	0.0XaaIle: 0.0 ± 0.0	0.0XaaLys: 0.0 ± 0.0	0.0XaaLeu: 0.0 ± 0.0	0.0XaaMet: 0.0 ± 0.0	0.0XaaAsn: 0.0 ± 0.0	0.0XaaPro: 0.0 ± 0.0	0.0XaaGln: 0.0 ± 0.0	0.0XaaArg: 0.0 ± 0.0	0.0XaaSer: 0.0 ± 0.0	0.0XaaThr: 0.0 ± 0.0	0.0XaaVal: 0.0 ± 0.0	0.0XaaTrp: 0.0 ± 0.0	0.0XaaTyr: 0.0 ± 0.0	0.0XaaXaa: 0.0 ± 0.0

Statistics based on 10363 proteins (5172183 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944

Contact: Lukasz P. Kozlowski

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