Amino acid dipepetide frequency for Bean golden yellow mosaic virus (isolate Puerto Rico-Japan) (BGYMV)

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
0.57AlaAla: 0.57 ± 0.46
1.14AlaCys: 1.14 ± 0.716
2.281AlaAsp: 2.281 ± 0.87
0.57AlaGlu: 0.57 ± 0.549
1.14AlaPhe: 1.14 ± 0.69
1.71AlaGly: 1.71 ± 1.131
1.14AlaHis: 1.14 ± 0.722
3.991AlaIle: 3.991 ± 1.327
3.991AlaLys: 3.991 ± 0.678
5.701AlaLeu: 5.701 ± 2.67
0.57AlaMet: 0.57 ± 0.539
2.281AlaAsn: 2.281 ± 0.563
1.71AlaPro: 1.71 ± 0.595
5.131AlaGln: 5.131 ± 1.6
3.421AlaArg: 3.421 ± 1.203
6.271AlaSer: 6.271 ± 2.122
2.281AlaThr: 2.281 ± 1.061
1.14AlaVal: 1.14 ± 0.812
0.57AlaTrp: 0.57 ± 0.461
0.57AlaTyr: 0.57 ± 0.549
0.0AlaXaa: 0.0 ± 0.0
Cys
1.71CysAla: 1.71 ± 2.133
0.57CysCys: 0.57 ± 0.711
0.0CysAsp: 0.0 ± 0.0
1.14CysGlu: 1.14 ± 0.567
0.57CysPhe: 0.57 ± 0.624
0.57CysGly: 0.57 ± 0.711
0.0CysHis: 0.0 ± 0.0
3.991CysIle: 3.991 ± 1.501
1.14CysLys: 1.14 ± 0.567
1.14CysLeu: 1.14 ± 0.724
0.57CysMet: 0.57 ± 0.466
1.71CysAsn: 1.71 ± 0.606
0.0CysPro: 0.0 ± 0.0
1.14CysGln: 1.14 ± 0.613
0.57CysArg: 0.57 ± 0.577
1.14CysSer: 1.14 ± 0.931
2.281CysThr: 2.281 ± 0.948
2.281CysVal: 2.281 ± 1.274
0.57CysTrp: 0.57 ± 0.577
0.0CysTyr: 0.0 ± 0.0
0.0CysXaa: 0.0 ± 0.0
Asp
1.14AspAla: 1.14 ± 0.633
1.14AspCys: 1.14 ± 0.732
2.851AspAsp: 2.851 ± 1.562
1.71AspGlu: 1.71 ± 0.657
3.421AspPhe: 3.421 ± 1.089
2.281AspGly: 2.281 ± 1.38
2.281AspHis: 2.281 ± 1.343
5.701AspIle: 5.701 ± 1.711
2.281AspLys: 2.281 ± 0.603
5.701AspLeu: 5.701 ± 1.048
0.57AspMet: 0.57 ± 0.624
2.281AspAsn: 2.281 ± 0.716
2.281AspPro: 2.281 ± 1.051
0.57AspGln: 0.57 ± 0.624
2.851AspArg: 2.851 ± 1.279
6.271AspSer: 6.271 ± 1.356
1.71AspThr: 1.71 ± 0.809
3.421AspVal: 3.421 ± 1.085
1.14AspTrp: 1.14 ± 0.92
2.281AspTyr: 2.281 ± 0.881
0.0AspXaa: 0.0 ± 0.0
Glu
1.14GluAla: 1.14 ± 0.567
1.14GluCys: 1.14 ± 0.73
1.14GluAsp: 1.14 ± 0.812
3.421GluGlu: 3.421 ± 1.847
0.57GluPhe: 0.57 ± 0.711
3.421GluGly: 3.421 ± 1.571
0.57GluHis: 0.57 ± 0.711
1.71GluIle: 1.71 ± 1.397
0.0GluLys: 0.0 ± 0.0
3.421GluLeu: 3.421 ± 1.278
0.0GluMet: 0.0 ± 0.0
4.561GluAsn: 4.561 ± 1.963
3.421GluPro: 3.421 ± 0.907
1.71GluGln: 1.71 ± 0.926
2.851GluArg: 2.851 ± 1.268
4.561GluSer: 4.561 ± 1.817
0.0GluThr: 0.0 ± 0.0
1.14GluVal: 1.14 ± 0.804
1.14GluTrp: 1.14 ± 0.613
2.851GluTyr: 2.851 ± 1.034
0.0GluXaa: 0.0 ± 0.0
Phe
1.14PheAla: 1.14 ± 0.69
0.57PheCys: 0.57 ± 0.461
2.281PheAsp: 2.281 ± 1.12
0.57PheGlu: 0.57 ± 0.46
1.14PhePhe: 1.14 ± 0.522
2.281PheGly: 2.281 ± 1.12
0.57PheHis: 0.57 ± 0.46
3.991PheIle: 3.991 ± 1.392
3.421PheLys: 3.421 ± 1.07
2.281PheLeu: 2.281 ± 1.122
0.0PheMet: 0.0 ± 0.0
3.421PheAsn: 3.421 ± 0.917
2.281PhePro: 2.281 ± 0.991
1.71PheGln: 1.71 ± 0.747
3.421PheArg: 3.421 ± 0.802
4.561PheSer: 4.561 ± 2.211
0.57PheThr: 0.57 ± 0.711
2.851PheVal: 2.851 ± 1.321
1.71PheTrp: 1.71 ± 1.131
2.851PheTyr: 2.851 ± 1.266
0.0PheXaa: 0.0 ± 0.0
Gly
2.281GlyAla: 2.281 ± 0.881
2.281GlyCys: 2.281 ± 0.947
1.71GlyAsp: 1.71 ± 0.691
2.851GlyGlu: 2.851 ± 0.895
1.14GlyPhe: 1.14 ± 0.77
3.421GlyGly: 3.421 ± 1.523
0.57GlyHis: 0.57 ± 0.46
1.71GlyIle: 1.71 ± 0.749
6.271GlyLys: 6.271 ± 1.721
1.71GlyLeu: 1.71 ± 0.809
2.281GlyMet: 2.281 ± 0.883
1.71GlyAsn: 1.71 ± 0.959
3.421GlyPro: 3.421 ± 0.66
2.851GlyGln: 2.851 ± 1.233
2.281GlyArg: 2.281 ± 1.424
5.701GlySer: 5.701 ± 0.726
2.851GlyThr: 2.851 ± 1.013
3.421GlyVal: 3.421 ± 1.173
0.0GlyTrp: 0.0 ± 0.0
1.14GlyTyr: 1.14 ± 0.724
0.0GlyXaa: 0.0 ± 0.0
His
1.71HisAla: 1.71 ± 0.762
1.14HisCys: 1.14 ± 0.818
4.561HisAsp: 4.561 ± 1.076
0.0HisGlu: 0.0 ± 0.0
1.14HisPhe: 1.14 ± 0.522
1.71HisGly: 1.71 ± 0.921
0.57HisHis: 0.57 ± 0.711
1.71HisIle: 1.71 ± 0.963
1.14HisLys: 1.14 ± 0.74
3.991HisLeu: 3.991 ± 1.465
1.71HisMet: 1.71 ± 0.824
3.991HisAsn: 3.991 ± 1.89
2.281HisPro: 2.281 ± 0.956
2.851HisGln: 2.851 ± 0.744
3.991HisArg: 3.991 ± 1.652
0.57HisSer: 0.57 ± 0.466
3.991HisThr: 3.991 ± 1.222
3.421HisVal: 3.421 ± 1.465
0.57HisTrp: 0.57 ± 0.46
0.57HisTyr: 0.57 ± 0.466
0.0HisXaa: 0.0 ± 0.0
Ile
1.71IleAla: 1.71 ± 0.766
0.57IleCys: 0.57 ± 0.46
5.131IleAsp: 5.131 ± 1.488
5.701IleGlu: 5.701 ± 1.599
2.851IlePhe: 2.851 ± 1.401
3.421IleGly: 3.421 ± 1.155
4.561IleHis: 4.561 ± 1.775
2.281IleIle: 2.281 ± 0.991
4.561IleLys: 4.561 ± 1.897
3.991IleLeu: 3.991 ± 1.476
2.281IleMet: 2.281 ± 0.794
2.851IleAsn: 2.851 ± 1.337
4.561IlePro: 4.561 ± 1.574
2.851IleGln: 2.851 ± 1.285
5.131IleArg: 5.131 ± 0.862
3.991IleSer: 3.991 ± 1.14
3.991IleThr: 3.991 ± 1.714
5.131IleVal: 5.131 ± 1.737
2.281IleTrp: 2.281 ± 0.938
2.281IleTyr: 2.281 ± 1.117
0.0IleXaa: 0.0 ± 0.0
Lys
4.561LysAla: 4.561 ± 1.288
0.57LysCys: 0.57 ± 0.711
6.271LysAsp: 6.271 ± 1.373
3.421LysGlu: 3.421 ± 2.178
2.281LysPhe: 2.281 ± 1.013
2.281LysGly: 2.281 ± 0.563
2.851LysHis: 2.851 ± 1.447
6.271LysIle: 6.271 ± 1.604
0.57LysLys: 0.57 ± 0.46
3.991LysLeu: 3.991 ± 1.574
2.281LysMet: 2.281 ± 0.891
4.561LysAsn: 4.561 ± 1.246
2.281LysPro: 2.281 ± 0.563
0.57LysGln: 0.57 ± 0.466
5.701LysArg: 5.701 ± 2.37
5.701LysSer: 5.701 ± 1.151
2.281LysThr: 2.281 ± 1.526
4.561LysVal: 4.561 ± 2.286
0.57LysTrp: 0.57 ± 0.466
2.851LysTyr: 2.851 ± 0.734
0.0LysXaa: 0.0 ± 0.0
Leu
2.281LeuAla: 2.281 ± 1.181
1.71LeuCys: 1.71 ± 0.804
4.561LeuAsp: 4.561 ± 1.221
1.14LeuGlu: 1.14 ± 0.722
2.281LeuPhe: 2.281 ± 0.934
3.421LeuGly: 3.421 ± 0.487
3.421LeuHis: 3.421 ± 1.408
2.281LeuIle: 2.281 ± 0.963
7.982LeuLys: 7.982 ± 1.254
4.561LeuLeu: 4.561 ± 1.956
0.57LeuMet: 0.57 ± 0.539
4.561LeuAsn: 4.561 ± 1.335
2.281LeuPro: 2.281 ± 1.277
3.991LeuGln: 3.991 ± 1.362
1.71LeuArg: 1.71 ± 0.818
7.982LeuSer: 7.982 ± 2.646
3.991LeuThr: 3.991 ± 2.051
6.271LeuVal: 6.271 ± 1.946
0.0LeuTrp: 0.0 ± 0.0
3.421LeuTyr: 3.421 ± 1.018
0.0LeuXaa: 0.0 ± 0.0
Met
1.14MetAla: 1.14 ± 0.921
1.14MetCys: 1.14 ± 0.712
2.851MetAsp: 2.851 ± 1.024
1.14MetGlu: 1.14 ± 0.931
0.57MetPhe: 0.57 ± 0.461
0.57MetGly: 0.57 ± 0.461
2.281MetHis: 2.281 ± 0.884
1.71MetIle: 1.71 ± 0.887
0.57MetLys: 0.57 ± 0.466
1.71MetLeu: 1.71 ± 0.781
0.0MetMet: 0.0 ± 0.0
0.57MetAsn: 0.57 ± 0.539
2.281MetPro: 2.281 ± 0.724
0.57MetGln: 0.57 ± 0.46
2.281MetArg: 2.281 ± 1.066
1.71MetSer: 1.71 ± 1.732
0.57MetThr: 0.57 ± 0.466
1.71MetVal: 1.71 ± 1.35
1.14MetTrp: 1.14 ± 0.69
1.71MetTyr: 1.71 ± 0.959
0.0MetXaa: 0.0 ± 0.0
Asn
5.131AsnAla: 5.131 ± 1.296
1.71AsnCys: 1.71 ± 0.718
2.281AsnAsp: 2.281 ± 0.731
3.421AsnGlu: 3.421 ± 1.479
0.57AsnPhe: 0.57 ± 0.549
2.281AsnGly: 2.281 ± 0.848
4.561AsnHis: 4.561 ± 1.979
4.561AsnIle: 4.561 ± 1.239
4.561AsnLys: 4.561 ± 1.453
3.421AsnLeu: 3.421 ± 1.269
2.281AsnMet: 2.281 ± 1.264
2.851AsnAsn: 2.851 ± 1.206
2.851AsnPro: 2.851 ± 0.828
1.14AsnGln: 1.14 ± 0.802
2.851AsnArg: 2.851 ± 0.734
3.991AsnSer: 3.991 ± 1.551
1.71AsnThr: 1.71 ± 0.994
2.281AsnVal: 2.281 ± 0.819
0.57AsnTrp: 0.57 ± 0.46
3.991AsnTyr: 3.991 ± 1.359
0.0AsnXaa: 0.0 ± 0.0
Pro
0.57ProAla: 0.57 ± 0.549
1.71ProCys: 1.71 ± 1.176
1.14ProAsp: 1.14 ± 0.643
2.281ProGlu: 2.281 ± 1.018
1.71ProPhe: 1.71 ± 0.718
2.281ProGly: 2.281 ± 0.923
2.281ProHis: 2.281 ± 0.939
4.561ProIle: 4.561 ± 2.366
5.131ProLys: 5.131 ± 1.282
2.281ProLeu: 2.281 ± 0.995
2.281ProMet: 2.281 ± 1.16
2.851ProAsn: 2.851 ± 1.038
3.421ProPro: 3.421 ± 1.216
2.851ProGln: 2.851 ± 1.403
1.71ProArg: 1.71 ± 0.701
5.701ProSer: 5.701 ± 2.441
2.281ProThr: 2.281 ± 1.192
2.851ProVal: 2.851 ± 1.006
1.71ProTrp: 1.71 ± 0.595
2.281ProTyr: 2.281 ± 1.334
0.0ProXaa: 0.0 ± 0.0
Gln
2.281GlnAla: 2.281 ± 1.174
0.57GlnCys: 0.57 ± 0.577
1.71GlnAsp: 1.71 ± 0.965
1.71GlnGlu: 1.71 ± 0.998
3.421GlnPhe: 3.421 ± 1.098
2.851GlnGly: 2.851 ± 0.783
1.71GlnHis: 1.71 ± 0.969
1.14GlnIle: 1.14 ± 0.732
1.14GlnLys: 1.14 ± 0.92
3.421GlnLeu: 3.421 ± 1.625
0.57GlnMet: 0.57 ± 0.539
1.14GlnAsn: 1.14 ± 0.613
3.421GlnPro: 3.421 ± 1.601
1.14GlnGln: 1.14 ± 0.522
3.991GlnArg: 3.991 ± 0.82
6.271GlnSer: 6.271 ± 1.945
0.57GlnThr: 0.57 ± 0.46
5.701GlnVal: 5.701 ± 1.539
0.0GlnTrp: 0.0 ± 0.0
0.57GlnTyr: 0.57 ± 0.461
0.0GlnXaa: 0.0 ± 0.0
Arg
2.851ArgAla: 2.851 ± 1.15
1.71ArgCys: 1.71 ± 0.606
2.851ArgAsp: 2.851 ± 1.54
2.281ArgGlu: 2.281 ± 1.053
7.412ArgPhe: 7.412 ± 1.329
4.561ArgGly: 4.561 ± 1.038
3.991ArgHis: 3.991 ± 1.267
3.991ArgIle: 3.991 ± 1.221
2.851ArgLys: 2.851 ± 0.744
3.991ArgLeu: 3.991 ± 1.292
1.71ArgMet: 1.71 ± 0.657
0.0ArgAsn: 0.0 ± 0.0
2.851ArgPro: 2.851 ± 1.156
1.14ArgGln: 1.14 ± 0.617
5.131ArgArg: 5.131 ± 2.316
6.842ArgSer: 6.842 ± 1.32
3.421ArgThr: 3.421 ± 1.336
3.991ArgVal: 3.991 ± 0.587
0.0ArgTrp: 0.0 ± 0.0
1.71ArgTyr: 1.71 ± 1.132
0.0ArgXaa: 0.0 ± 0.0
Ser
5.701SerAla: 5.701 ± 2.34
2.281SerCys: 2.281 ± 0.988
2.851SerAsp: 2.851 ± 0.782
0.0SerGlu: 0.0 ± 0.0
1.71SerPhe: 1.71 ± 0.718
3.991SerGly: 3.991 ± 1.218
4.561SerHis: 4.561 ± 1.864
7.982SerIle: 7.982 ± 2.224
6.271SerLys: 6.271 ± 1.816
3.991SerLeu: 3.991 ± 1.743
0.0SerMet: 0.0 ± 0.0
7.982SerAsn: 7.982 ± 1.898
5.131SerPro: 5.131 ± 1.978
4.561SerGln: 4.561 ± 2.211
3.991SerArg: 3.991 ± 0.587
9.692SerSer: 9.692 ± 3.495
10.832SerThr: 10.832 ± 2.17
5.131SerVal: 5.131 ± 1.348
1.14SerTrp: 1.14 ± 0.931
3.421SerTyr: 3.421 ± 1.395
0.0SerXaa: 0.0 ± 0.0
Thr
5.131ThrAla: 5.131 ± 1.182
0.0ThrCys: 0.0 ± 0.0
2.281ThrAsp: 2.281 ± 0.988
1.71ThrGlu: 1.71 ± 0.903
3.421ThrPhe: 3.421 ± 2.24
3.421ThrGly: 3.421 ± 1.333
2.851ThrHis: 2.851 ± 1.239
3.991ThrIle: 3.991 ± 1.92
3.421ThrLys: 3.421 ± 2.18
2.281ThrLeu: 2.281 ± 0.563
1.71ThrMet: 1.71 ± 1.034
3.421ThrAsn: 3.421 ± 1.126
1.14ThrPro: 1.14 ± 0.716
0.57ThrGln: 0.57 ± 0.57
4.561ThrArg: 4.561 ± 1.453
2.851ThrSer: 2.851 ± 1.404
5.131ThrThr: 5.131 ± 1.655
3.421ThrVal: 3.421 ± 1.075
0.57ThrTrp: 0.57 ± 0.57
1.14ThrTyr: 1.14 ± 0.722
0.0ThrXaa: 0.0 ± 0.0
Val
0.57ValAla: 0.57 ± 0.624
0.57ValCys: 0.57 ± 0.46
3.421ValAsp: 3.421 ± 1.212
3.421ValGlu: 3.421 ± 0.991
3.421ValPhe: 3.421 ± 1.327
2.281ValGly: 2.281 ± 1.058
1.14ValHis: 1.14 ± 0.778
5.131ValIle: 5.131 ± 0.926
4.561ValLys: 4.561 ± 1.234
6.271ValLeu: 6.271 ± 1.936
3.991ValMet: 3.991 ± 1.404
4.561ValAsn: 4.561 ± 0.955
3.991ValPro: 3.991 ± 0.941
3.991ValGln: 3.991 ± 0.587
3.421ValArg: 3.421 ± 1.254
4.561ValSer: 4.561 ± 0.988
2.281ValThr: 2.281 ± 1.052
4.561ValVal: 4.561 ± 1.282
0.57ValTrp: 0.57 ± 0.549
3.991ValTyr: 3.991 ± 1.484
0.0ValXaa: 0.0 ± 0.0
Trp
1.71TrpAla: 1.71 ± 0.68
0.0TrpCys: 0.0 ± 0.0
0.57TrpAsp: 0.57 ± 0.711
1.14TrpGlu: 1.14 ± 0.69
0.0TrpPhe: 0.0 ± 0.0
0.57TrpGly: 0.57 ± 0.46
0.0TrpHis: 0.0 ± 0.0
0.0TrpIle: 0.0 ± 0.0
1.71TrpLys: 1.71 ± 0.595
0.57TrpLeu: 0.57 ± 0.461
1.14TrpMet: 1.14 ± 0.643
0.0TrpAsn: 0.0 ± 0.0
0.0TrpPro: 0.0 ± 0.0
0.57TrpGln: 0.57 ± 0.46
1.14TrpArg: 1.14 ± 0.712
1.14TrpSer: 1.14 ± 0.78
1.71TrpThr: 1.71 ± 0.735
1.71TrpVal: 1.71 ± 0.924
0.0TrpTrp: 0.0 ± 0.0
0.57TrpTyr: 0.57 ± 0.57
0.0TrpXaa: 0.0 ± 0.0
Tyr
2.851TyrAla: 2.851 ± 1.235
0.57TyrCys: 0.57 ± 0.577
1.14TyrAsp: 1.14 ± 0.643
1.14TyrGlu: 1.14 ± 0.921
2.851TyrPhe: 2.851 ± 0.967
2.281TyrGly: 2.281 ± 0.905
1.71TyrHis: 1.71 ± 1.343
3.421TyrIle: 3.421 ± 0.971
2.851TyrLys: 2.851 ± 0.734
3.991TyrLeu: 3.991 ± 1.322
1.14TyrMet: 1.14 ± 0.651
1.71TyrAsn: 1.71 ± 0.674
2.281TyrPro: 2.281 ± 0.875
3.421TyrGln: 3.421 ± 0.955
2.281TyrArg: 2.281 ± 1.265
1.71TyrSer: 1.71 ± 0.674
1.14TyrThr: 1.14 ± 0.74
1.71TyrVal: 1.71 ± 0.9
0.0TyrTrp: 0.0 ± 0.0
1.14TyrTyr: 1.14 ± 0.617
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 9 proteins (1755 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski