Amino acid dipepetide frequency for Tortoise microvirus 39

Apart from single amino acid frequencies one can also calculate so called amino acid dipeptide frequency.

There is 400 possibilites (441 if we consider Xaa as additional 21st amino acid representing all non-standard or unknown amino acids). Thus, if the dipeptides would be present randomly in proteins each amino acid dipeptide should be present with 0.25% frequency. As it is not the case in the nature, for better visablity all more than expected dipepetides are marked by red, and those which are underrepresented are marked by blue in the table.

All values are presented as per milles (‰), therefore need to be multiplied by 10-3.

For more information see sequence space article on Wikipedia.

AlaCysAspGluPheGlyHisIleLysLeuMetAsnProGlnArgSerThrValTrpTyrXaa
Ala
10.49AlaAla: 10.49 ± 2.476
1.748AlaCys: 1.748 ± 1.479
4.662AlaAsp: 4.662 ± 1.769
2.914AlaGlu: 2.914 ± 0.965
6.41AlaPhe: 6.41 ± 2.109
10.49AlaGly: 10.49 ± 3.772
5.245AlaHis: 5.245 ± 2.175
1.748AlaIle: 1.748 ± 0.755
6.41AlaLys: 6.41 ± 2.049
6.41AlaLeu: 6.41 ± 1.87
4.079AlaMet: 4.079 ± 1.066
2.914AlaAsn: 2.914 ± 1.338
5.828AlaPro: 5.828 ± 1.194
4.079AlaGln: 4.079 ± 0.922
4.079AlaArg: 4.079 ± 1.612
2.914AlaSer: 2.914 ± 1.119
5.828AlaThr: 5.828 ± 2.019
9.324AlaVal: 9.324 ± 2.1
2.914AlaTrp: 2.914 ± 1.448
3.497AlaTyr: 3.497 ± 1.129
0.0AlaXaa: 0.0 ± 0.0
Cys
0.0CysAla: 0.0 ± 0.0
0.0CysCys: 0.0 ± 0.0
0.0CysAsp: 0.0 ± 0.0
0.583CysGlu: 0.583 ± 0.411
0.0CysPhe: 0.0 ± 0.0
0.583CysGly: 0.583 ± 0.493
0.0CysHis: 0.0 ± 0.0
1.166CysIle: 1.166 ± 0.986
0.583CysLys: 0.583 ± 0.493
0.0CysLeu: 0.0 ± 0.0
0.0CysMet: 0.0 ± 0.0
0.583CysAsn: 0.583 ± 0.493
0.0CysPro: 0.0 ± 0.0
1.166CysGln: 1.166 ± 0.986
1.748CysArg: 1.748 ± 0.891
0.0CysSer: 0.0 ± 0.0
0.583CysThr: 0.583 ± 0.411
0.583CysVal: 0.583 ± 0.628
0.583CysTrp: 0.583 ± 0.493
0.0CysTyr: 0.0 ± 0.0
0.0CysXaa: 0.0 ± 0.0
Asp
3.497AspAla: 3.497 ± 1.705
0.0AspCys: 0.0 ± 0.0
3.497AspAsp: 3.497 ± 0.877
2.331AspGlu: 2.331 ± 1.114
2.914AspPhe: 2.914 ± 0.564
6.993AspGly: 6.993 ± 3.618
1.748AspHis: 1.748 ± 0.644
1.748AspIle: 1.748 ± 1.329
1.166AspLys: 1.166 ± 0.791
6.41AspLeu: 6.41 ± 1.97
0.583AspMet: 0.583 ± 0.411
1.166AspAsn: 1.166 ± 0.927
6.993AspPro: 6.993 ± 1.236
2.331AspGln: 2.331 ± 0.844
5.828AspArg: 5.828 ± 2.566
0.583AspSer: 0.583 ± 0.411
2.331AspThr: 2.331 ± 0.857
3.497AspVal: 3.497 ± 1.667
1.748AspTrp: 1.748 ± 1.334
1.166AspTyr: 1.166 ± 0.568
0.0AspXaa: 0.0 ± 0.0
Glu
8.159GluAla: 8.159 ± 1.948
0.583GluCys: 0.583 ± 0.493
2.914GluAsp: 2.914 ± 1.059
2.914GluGlu: 2.914 ± 1.811
2.331GluPhe: 2.331 ± 1.238
4.079GluGly: 4.079 ± 1.584
0.583GluHis: 0.583 ± 0.411
1.748GluIle: 1.748 ± 0.81
1.748GluLys: 1.748 ± 0.953
5.828GluLeu: 5.828 ± 2.014
1.166GluMet: 1.166 ± 0.486
1.166GluAsn: 1.166 ± 0.986
0.583GluPro: 0.583 ± 0.646
2.914GluGln: 2.914 ± 0.725
4.079GluArg: 4.079 ± 0.694
2.331GluSer: 2.331 ± 1.063
3.497GluThr: 3.497 ± 0.749
5.828GluVal: 5.828 ± 2.117
1.166GluTrp: 1.166 ± 0.632
2.331GluTyr: 2.331 ± 0.976
0.0GluXaa: 0.0 ± 0.0
Phe
2.331PheAla: 2.331 ± 1.193
0.583PheCys: 0.583 ± 0.493
2.914PheAsp: 2.914 ± 0.943
3.497PheGlu: 3.497 ± 1.948
1.166PhePhe: 1.166 ± 0.488
4.079PheGly: 4.079 ± 1.051
0.0PheHis: 0.0 ± 0.0
1.748PheIle: 1.748 ± 1.31
1.166PheLys: 1.166 ± 0.986
3.497PheLeu: 3.497 ± 0.894
1.166PheMet: 1.166 ± 0.835
1.166PheAsn: 1.166 ± 0.568
1.166PhePro: 1.166 ± 0.673
1.166PheGln: 1.166 ± 1.144
2.331PheArg: 2.331 ± 1.355
0.583PheSer: 0.583 ± 0.493
1.748PheThr: 1.748 ± 1.063
3.497PheVal: 3.497 ± 0.96
1.748PheTrp: 1.748 ± 0.891
2.914PheTyr: 2.914 ± 1.205
0.0PheXaa: 0.0 ± 0.0
Gly
5.828GlyAla: 5.828 ± 1.948
0.0GlyCys: 0.0 ± 0.0
2.914GlyAsp: 2.914 ± 1.326
8.159GlyGlu: 8.159 ± 1.876
1.748GlyPhe: 1.748 ± 1.147
11.072GlyGly: 11.072 ± 2.183
2.331GlyHis: 2.331 ± 0.568
2.331GlyIle: 2.331 ± 1.072
2.914GlyLys: 2.914 ± 1.205
6.41GlyLeu: 6.41 ± 3.279
1.166GlyMet: 1.166 ± 0.822
2.914GlyAsn: 2.914 ± 1.068
1.748GlyPro: 1.748 ± 1.063
2.331GlyGln: 2.331 ± 1.057
9.324GlyArg: 9.324 ± 2.849
4.079GlySer: 4.079 ± 1.081
4.662GlyThr: 4.662 ± 1.596
6.993GlyVal: 6.993 ± 2.258
0.0GlyTrp: 0.0 ± 0.0
1.166GlyTyr: 1.166 ± 0.488
0.0GlyXaa: 0.0 ± 0.0
His
2.331HisAla: 2.331 ± 0.844
1.166HisCys: 1.166 ± 0.632
2.331HisAsp: 2.331 ± 0.798
1.166HisGlu: 1.166 ± 0.835
1.748HisPhe: 1.748 ± 0.889
4.662HisGly: 4.662 ± 1.982
0.583HisHis: 0.583 ± 0.493
0.0HisIle: 0.0 ± 0.0
0.583HisLys: 0.583 ± 0.628
0.0HisLeu: 0.0 ± 0.0
0.583HisMet: 0.583 ± 0.572
1.748HisAsn: 1.748 ± 0.816
1.748HisPro: 1.748 ± 1.151
0.583HisGln: 0.583 ± 0.411
0.583HisArg: 0.583 ± 0.572
2.331HisSer: 2.331 ± 1.264
1.748HisThr: 1.748 ± 0.524
2.331HisVal: 2.331 ± 0.568
1.748HisTrp: 1.748 ± 1.334
0.583HisTyr: 0.583 ± 0.628
0.0HisXaa: 0.0 ± 0.0
Ile
1.748IleAla: 1.748 ± 1.01
0.583IleCys: 0.583 ± 0.493
2.331IleAsp: 2.331 ± 1.267
2.914IleGlu: 2.914 ± 1.24
1.748IlePhe: 1.748 ± 0.891
1.166IleGly: 1.166 ± 0.568
0.0IleHis: 0.0 ± 0.0
1.166IleIle: 1.166 ± 0.822
0.0IleLys: 0.0 ± 0.0
2.914IleLeu: 2.914 ± 1.448
0.583IleMet: 0.583 ± 0.572
1.166IleAsn: 1.166 ± 0.488
2.914IlePro: 2.914 ± 1.1
0.0IleGln: 0.0 ± 0.0
2.331IleArg: 2.331 ± 1.442
4.079IleSer: 4.079 ± 1.222
4.079IleThr: 4.079 ± 1.297
2.914IleVal: 2.914 ± 1.343
0.583IleTrp: 0.583 ± 0.411
0.583IleTyr: 0.583 ± 0.646
0.0IleXaa: 0.0 ± 0.0
Lys
2.914LysAla: 2.914 ± 0.517
0.0LysCys: 0.0 ± 0.0
2.331LysAsp: 2.331 ± 0.659
1.748LysGlu: 1.748 ± 1.187
1.748LysPhe: 1.748 ± 0.891
1.748LysGly: 1.748 ± 0.816
0.583LysHis: 0.583 ± 0.411
1.166LysIle: 1.166 ± 1.055
1.748LysLys: 1.748 ± 1.479
5.245LysLeu: 5.245 ± 1.805
0.583LysMet: 0.583 ± 0.493
1.748LysAsn: 1.748 ± 0.81
4.079LysPro: 4.079 ± 1.206
2.914LysGln: 2.914 ± 1.032
2.331LysArg: 2.331 ± 0.568
1.748LysSer: 1.748 ± 1.328
0.583LysThr: 0.583 ± 0.646
2.914LysVal: 2.914 ± 1.034
1.166LysTrp: 1.166 ± 0.791
1.748LysTyr: 1.748 ± 0.816
0.0LysXaa: 0.0 ± 0.0
Leu
13.403LeuAla: 13.403 ± 3.062
0.583LeuCys: 0.583 ± 0.411
2.331LeuAsp: 2.331 ± 0.976
5.828LeuGlu: 5.828 ± 2.891
1.748LeuPhe: 1.748 ± 0.81
5.245LeuGly: 5.245 ± 2.687
1.748LeuHis: 1.748 ± 0.861
1.748LeuIle: 1.748 ± 1.765
2.331LeuLys: 2.331 ± 1.136
8.159LeuLeu: 8.159 ± 1.646
2.331LeuMet: 2.331 ± 1.201
4.662LeuAsn: 4.662 ± 1.291
2.331LeuPro: 2.331 ± 0.976
1.748LeuGln: 1.748 ± 0.715
5.828LeuArg: 5.828 ± 1.718
4.079LeuSer: 4.079 ± 1.508
7.576LeuThr: 7.576 ± 1.119
4.662LeuVal: 4.662 ± 1.098
0.583LeuTrp: 0.583 ± 0.744
2.331LeuTyr: 2.331 ± 1.291
0.0LeuXaa: 0.0 ± 0.0
Met
1.748MetAla: 1.748 ± 0.838
0.583MetCys: 0.583 ± 0.493
1.748MetAsp: 1.748 ± 0.81
1.748MetGlu: 1.748 ± 1.181
0.0MetPhe: 0.0 ± 0.0
1.166MetGly: 1.166 ± 0.488
1.748MetHis: 1.748 ± 0.669
1.748MetIle: 1.748 ± 1.118
2.331MetLys: 2.331 ± 0.908
4.079MetLeu: 4.079 ± 1.606
1.166MetMet: 1.166 ± 1.144
2.331MetAsn: 2.331 ± 1.136
0.583MetPro: 0.583 ± 0.493
1.166MetGln: 1.166 ± 0.791
2.331MetArg: 2.331 ± 0.785
3.497MetSer: 3.497 ± 1.193
0.583MetThr: 0.583 ± 0.572
0.583MetVal: 0.583 ± 0.411
0.0MetTrp: 0.0 ± 0.0
0.583MetTyr: 0.583 ± 0.572
0.0MetXaa: 0.0 ± 0.0
Asn
4.662AsnAla: 4.662 ± 0.663
0.0AsnCys: 0.0 ± 0.0
1.166AsnAsp: 1.166 ± 0.568
1.748AsnGlu: 1.748 ± 0.788
1.166AsnPhe: 1.166 ± 0.822
4.079AsnGly: 4.079 ± 0.515
1.748AsnHis: 1.748 ± 0.626
3.497AsnIle: 3.497 ± 0.807
1.748AsnLys: 1.748 ± 1.069
1.748AsnLeu: 1.748 ± 0.934
1.166AsnMet: 1.166 ± 1.075
1.748AsnAsn: 1.748 ± 0.769
3.497AsnPro: 3.497 ± 1.294
0.0AsnGln: 0.0 ± 0.0
5.828AsnArg: 5.828 ± 2.36
3.497AsnSer: 3.497 ± 1.329
2.331AsnThr: 2.331 ± 1.072
4.079AsnVal: 4.079 ± 1.835
0.583AsnTrp: 0.583 ± 0.646
1.748AsnTyr: 1.748 ± 0.818
0.0AsnXaa: 0.0 ± 0.0
Pro
4.662ProAla: 4.662 ± 1.652
1.166ProCys: 1.166 ± 0.488
4.079ProAsp: 4.079 ± 1.402
4.079ProGlu: 4.079 ± 1.478
4.079ProPhe: 4.079 ± 1.852
3.497ProGly: 3.497 ± 0.39
1.748ProHis: 1.748 ± 1.151
2.331ProIle: 2.331 ± 0.659
0.583ProLys: 0.583 ± 0.572
5.245ProLeu: 5.245 ± 1.724
0.583ProMet: 0.583 ± 0.411
2.331ProAsn: 2.331 ± 0.844
0.583ProPro: 0.583 ± 0.493
3.497ProGln: 3.497 ± 1.338
1.748ProArg: 1.748 ± 0.769
1.748ProSer: 1.748 ± 0.889
5.245ProThr: 5.245 ± 1.077
6.993ProVal: 6.993 ± 1.985
0.583ProTrp: 0.583 ± 0.493
1.166ProTyr: 1.166 ± 0.986
0.0ProXaa: 0.0 ± 0.0
Gln
5.245GlnAla: 5.245 ± 1.013
0.583GlnCys: 0.583 ± 0.493
2.331GlnAsp: 2.331 ± 0.659
0.0GlnGlu: 0.0 ± 0.0
0.583GlnPhe: 0.583 ± 0.572
3.497GlnGly: 3.497 ± 1.81
1.166GlnHis: 1.166 ± 0.786
0.583GlnIle: 0.583 ± 0.411
1.748GlnLys: 1.748 ± 0.755
4.079GlnLeu: 4.079 ± 0.725
2.331GlnMet: 2.331 ± 1.049
2.914GlnAsn: 2.914 ± 0.725
1.166GlnPro: 1.166 ± 0.488
1.748GlnGln: 1.748 ± 0.524
5.828GlnArg: 5.828 ± 2.632
1.748GlnSer: 1.748 ± 1.479
1.748GlnThr: 1.748 ± 1.069
2.914GlnVal: 2.914 ± 0.942
0.583GlnTrp: 0.583 ± 0.493
1.748GlnTyr: 1.748 ± 0.818
0.0GlnXaa: 0.0 ± 0.0
Arg
8.741ArgAla: 8.741 ± 2.322
0.583ArgCys: 0.583 ± 0.493
5.245ArgAsp: 5.245 ± 1.389
3.497ArgGlu: 3.497 ± 1.086
4.662ArgPhe: 4.662 ± 2.099
1.748ArgGly: 1.748 ± 1.191
4.079ArgHis: 4.079 ± 1.562
2.331ArgIle: 2.331 ± 0.798
4.079ArgLys: 4.079 ± 1.67
4.079ArgLeu: 4.079 ± 1.298
3.497ArgMet: 3.497 ± 1.226
4.079ArgAsn: 4.079 ± 2.247
2.914ArgPro: 2.914 ± 0.988
2.914ArgGln: 2.914 ± 0.887
5.828ArgArg: 5.828 ± 2.301
2.914ArgSer: 2.914 ± 1.349
6.41ArgThr: 6.41 ± 0.65
2.331ArgVal: 2.331 ± 1.136
0.583ArgTrp: 0.583 ± 0.572
2.914ArgTyr: 2.914 ± 1.065
0.0ArgXaa: 0.0 ± 0.0
Ser
5.828SerAla: 5.828 ± 0.975
0.0SerCys: 0.0 ± 0.0
2.914SerAsp: 2.914 ± 0.754
2.914SerGlu: 2.914 ± 0.618
1.166SerPhe: 1.166 ± 0.488
2.331SerGly: 2.331 ± 0.857
1.748SerHis: 1.748 ± 1.032
0.583SerIle: 0.583 ± 0.493
2.331SerLys: 2.331 ± 0.892
2.914SerLeu: 2.914 ± 1.274
2.914SerMet: 2.914 ± 0.952
1.748SerAsn: 1.748 ± 1.001
4.662SerPro: 4.662 ± 1.103
1.748SerGln: 1.748 ± 1.032
0.0SerArg: 0.0 ± 0.0
1.748SerSer: 1.748 ± 0.715
2.914SerThr: 2.914 ± 1.018
4.079SerVal: 4.079 ± 2.297
0.583SerTrp: 0.583 ± 0.572
2.331SerTyr: 2.331 ± 1.355
0.0SerXaa: 0.0 ± 0.0
Thr
5.828ThrAla: 5.828 ± 1.042
0.0ThrCys: 0.0 ± 0.0
2.914ThrAsp: 2.914 ± 1.936
2.331ThrGlu: 2.331 ± 1.194
1.166ThrPhe: 1.166 ± 0.632
6.41ThrGly: 6.41 ± 1.511
0.0ThrHis: 0.0 ± 0.0
2.914ThrIle: 2.914 ± 1.138
2.331ThrLys: 2.331 ± 1.456
4.662ThrLeu: 4.662 ± 1.1
2.331ThrMet: 2.331 ± 0.749
2.914ThrAsn: 2.914 ± 1.328
4.662ThrPro: 4.662 ± 1.245
4.079ThrGln: 4.079 ± 1.381
3.497ThrArg: 3.497 ± 1.243
2.914ThrSer: 2.914 ± 1.338
2.914ThrThr: 2.914 ± 2.17
4.662ThrVal: 4.662 ± 1.786
0.583ThrTrp: 0.583 ± 0.572
3.497ThrTyr: 3.497 ± 1.745
0.0ThrXaa: 0.0 ± 0.0
Val
8.741ValAla: 8.741 ± 2.471
0.0ValCys: 0.0 ± 0.0
5.828ValAsp: 5.828 ± 1.449
4.662ValGlu: 4.662 ± 1.393
2.331ValPhe: 2.331 ± 1.203
3.497ValGly: 3.497 ± 2.123
1.748ValHis: 1.748 ± 0.889
2.331ValIle: 2.331 ± 1.114
4.079ValLys: 4.079 ± 1.425
2.914ValLeu: 2.914 ± 1.169
1.166ValMet: 1.166 ± 0.822
5.828ValAsn: 5.828 ± 2.569
7.576ValPro: 7.576 ± 3.709
2.914ValGln: 2.914 ± 0.975
6.993ValArg: 6.993 ± 2.326
2.914ValSer: 2.914 ± 0.965
4.662ValThr: 4.662 ± 1.329
4.662ValVal: 4.662 ± 1.788
0.583ValTrp: 0.583 ± 0.411
1.166ValTyr: 1.166 ± 0.69
0.0ValXaa: 0.0 ± 0.0
Trp
1.166TrpAla: 1.166 ± 0.822
0.0TrpCys: 0.0 ± 0.0
2.331TrpAsp: 2.331 ± 1.591
0.583TrpGlu: 0.583 ± 0.628
1.166TrpPhe: 1.166 ± 0.673
0.583TrpGly: 0.583 ± 0.628
1.166TrpHis: 1.166 ± 0.786
0.583TrpIle: 0.583 ± 0.572
1.166TrpLys: 1.166 ± 0.568
2.331TrpLeu: 2.331 ± 0.669
0.583TrpMet: 0.583 ± 0.493
0.583TrpAsn: 0.583 ± 0.411
0.583TrpPro: 0.583 ± 0.493
2.331TrpGln: 2.331 ± 1.488
1.166TrpArg: 1.166 ± 1.256
1.166TrpSer: 1.166 ± 0.728
0.0TrpThr: 0.0 ± 0.0
0.583TrpVal: 0.583 ± 0.493
0.583TrpTrp: 0.583 ± 0.572
0.0TrpTyr: 0.0 ± 0.0
0.0TrpXaa: 0.0 ± 0.0
Tyr
5.245TyrAla: 5.245 ± 1.053
0.0TyrCys: 0.0 ± 0.0
1.748TyrAsp: 1.748 ± 0.816
2.331TyrGlu: 2.331 ± 0.97
0.583TyrPhe: 0.583 ± 0.493
1.166TyrGly: 1.166 ± 0.986
0.583TyrHis: 0.583 ± 0.411
2.331TyrIle: 2.331 ± 0.976
0.0TyrLys: 0.0 ± 0.0
2.331TyrLeu: 2.331 ± 0.785
1.166TyrMet: 1.166 ± 0.488
2.331TyrAsn: 2.331 ± 1.096
2.331TyrPro: 2.331 ± 1.588
2.914TyrGln: 2.914 ± 1.032
1.748TyrArg: 1.748 ± 0.818
0.583TyrSer: 0.583 ± 0.493
1.166TyrThr: 1.166 ± 1.033
1.166TyrVal: 1.166 ± 0.488
1.748TyrTrp: 1.748 ± 1.479
1.748TyrTyr: 1.748 ± 1.479
0.0TyrXaa: 0.0 ± 0.0
Xaa
0.0XaaAla: 0.0 ± 0.0
0.0XaaCys: 0.0 ± 0.0
0.0XaaAsp: 0.0 ± 0.0
0.0XaaGlu: 0.0 ± 0.0
0.0XaaPhe: 0.0 ± 0.0
0.0XaaGly: 0.0 ± 0.0
0.0XaaHis: 0.0 ± 0.0
0.0XaaIle: 0.0 ± 0.0
0.0XaaLys: 0.0 ± 0.0
0.0XaaLeu: 0.0 ± 0.0
0.0XaaMet: 0.0 ± 0.0
0.0XaaAsn: 0.0 ± 0.0
0.0XaaPro: 0.0 ± 0.0
0.0XaaGln: 0.0 ± 0.0
0.0XaaArg: 0.0 ± 0.0
0.0XaaSer: 0.0 ± 0.0
0.0XaaThr: 0.0 ± 0.0
0.0XaaVal: 0.0 ± 0.0
0.0XaaTrp: 0.0 ± 0.0
0.0XaaTyr: 0.0 ± 0.0
0.0XaaXaa: 0.0 ± 0.0
Statistics based on 7 proteins (1717 amino acids)

Note: The error has been estimated with the bootstraping (x100) at the protein level

Above dipeptide statistics (among other stats for this proteome) you can download from this CSV file
See this proteome in: uniprot_link
Proteome-pI is available under Creative Commons Attribution-NoDerivs license, for more details see here

Reference: Kozlowski LP. Proteome-pI 2.0: Proteome Isoelectric Point Database Update. Nucleic Acids Res. 2021, doi: 10.1093/nar/gkab944 Contact: Lukasz P. Kozlowski